The ESEM investigations revealed the morphogenesis of peltate glandular trichomes, which was divided into five phases. In phase one, new peltate glandular trichomes were initiated; in phase two, they were differentiated; in phase three trichomes grew vigorously; in phase four they were determined; in the fifth and final phase they came to maturity. Volume of glandular trichome during the different phases of morphogenesis varied from 0.25´10 -2 mm 3 in phase 1, to 1.95´10 -2 mm 3 in phase 5. More glandular trichomes are placed on the base of the adaxial side of bracts (average 7 mm -2 ) than on the base of abaxial side (average 5.8 mm -2 ). In this research, positive Spearman's rank order correlations were found between the average number of glandular trichomes and content of a-acids as well as between the average volume of glandular trichomes and content of a-acids.
The results of this study show that the most usual damages of hop glandular trichomes are obtained on tunica of peltate glandular trichomes as well as cracking of bulbous glandular trichomes. In fresh hop cones any sorts of damage of glandular trichomes are obtained, on the other hand frequency of damaged hop glandular trichomes increase after drying (f i =0.08) and after pelletization of hop cones the structure of glandular trichomes is completely destroyed. Volume of hop glandular trichomes of dried hop cones is significantly smaller in the comparison with volume of glandular trichomes of fresh hop cones (D=0.544×10-2 mm 3 ; P<0.01) and a phenomenon of shrinkage on tunica surface of peltate glandular trichomes of dried hop cones is also visible in comparison with the fresh one. The decrease in volume of hop glandular trichomes in dried hop cones is the result of dehydration of hop cones and consequently hops glandular trichomes during the drying process which cause shrinkage of peltate glandular trichomes. However, after pelletization the structure of hop glandular trichomes is completely destroyed.
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