water use efficiency, stomatal activity, and osmotic adjustment), and biochemical reactions (accumulation of proline, polyamines, trehalose, increase of nitrate reductase activity, and storage of saccharides) (Haworth et al. 2013, Ammar et al. 2014, Conesa et al. 2016. In response to water stress, abscisic acid (ABA) accumulation is one of the key adaptive mechanisms (Pennisi 2017, Phillips and Sussman 2019).The rapid development of molecular biology in recent years has promoted research on the regulation mechanisms of plant responses to the environment at
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