Granular materials avalanche when a static angle of repose is exceeded and freeze at a dynamic angle of repose. Such avalanches occur subaerially on steep hillslopes and wind dunes and subaqueously at the lee side of deltas. Until now it has been assumed that the angles of repose are independent of gravitational acceleration. The objective of this work is to experimentally determine whether the angles of repose depend on gravity. In 33 parabolic flights in a well‐controlled research aircraft we recorded avalanching granular materials in rotating drums at effective gravitational accelerations of 0.1, 0.38 and 1.0 times the terrestrial value. The granular materials varied in particle size and rounding and had air or water as interstitial fluid. Materials with angular grains had time‐averaged angles of about 40° and with rounded grains about 25° for all effective gravitational accelerations, except the finest glass beads in air, which was explained by static electricity. For all materials, the static angle of repose increases about 5° with reduced gravity, whereas the dynamic angle decreases with about 10°. Consequently, the avalanche size increases with reduced gravity. The experimental results suggest that relatively low slopes of granular material on Mars may have formed by dry flows without a lubricating fluid. On asteroids even lower slopes are expected. The dependence on gravity of angle of repose may require reanalysis of models for many phenomena involving sediment, also at much lower slope angles.
Successful dispersal of freshwater sponges depends on the formation of dormant sponge bodies (gemmules) under adverse conditions. Gemmule formation allows the sponge to overcome critical environmental conditions, for example, desiccation or freezing, and to re-establish as a fully developed sponge when conditions are more favorable. A key process in sponge development from hatched gemmules is the construction of the silica skeleton. Silica spicules form the structural support for the three-dimensional filtration system the sponge uses to filter food particles from ambient water. We studied the effect of different hypergravity forces (1, 2.5, 5, 10, and 20 × g for 48 h)—as measure for environmental stress—on the ability of developing sponges to set-up their spiculous skeleton. Additionally, we assessed whether the addition of nutrients (i.e., dissolved 13C- and 15N-labeled amino acids) compensates for this stress. Our results show that freshwater sponges can withstand prolonged periods of hypergravity exposure and successfully set-up their skeleton, even after 48 h under 20 × g. Developing sponges were found to take up and assimilate dissolved food before forming a functional filtering system. However, fed and non-fed sponges showed no differences in skeleton formation and relative surface area growth, suggesting that the gemmules’ intrinsic energy fulfills the processes of skeleton construction. Additionally, non-fed sponges formed oscula significantly more often than fed sponges, especially under higher g-forces. This suggests that the eventual formation of a filtration system might be stimulated by food deprivation and environmentally stressful conditions. These findings indicate that the process of spiculous skeleton formation is energy-efficient and highly resilient. The uptake of dissolved food substances by freshwater sponges may contribute to the cycling of dissolved organic matter in freshwater ecosystems where sponges are abundant.
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