Expcrimcnts conducted with natural mixed populations of 800 and 1,800 tubificids m ' in sediment from Mcssalonskec Lake, Maine, showed average sediment transport by alimentation at 10°C 2-3 times greater than highest rates previously reported.Marc than QS% of feeding on introduced pollen was at depths above 7 cm, with greatest feeding at 34 cm. Small amounts of pollen were raised to the surface from as deep as 15 cm. Downward transport was 14 and 19% of upward.Small pollen grains ( < 40 ,u) were fed upon and displaced at higher rates than large grains. Organic matter was less in the surface layer of fcccs than in sediment from feeding depths and in smface scdimcnt whcrc no worms were present. A mathematical model was used to appraise the stratigraphic cffccts of the worms by deriving age-frequency composition of sediment at various depths.

The landscapes of northern New England and adjacent areas of Canada changed greatly between 14,000 and 9000 yr B.P.: deglaciation occurred, sea levels and shorelines shifted, and a vegetational transition from tundra to closed forest took place. Data from 51 14C-dated sites from a range of elevations were used to map ice and sea positions, physiognomic vegetational zones, and the spread of individual tree taxa in the region. A continuum of tundra-woodland-forest passed northeastward and northward without major hesitation or reversal. An increased rate of progression from 11,000 to 10,000 yr B.P. suggests a more rapid warming than in the prior 2000–3000 yr. Elevational gradients controlled the patterns of deglaciation and vegetational change. The earliest spread of tree taxa was via the lowlands of southern Vermont and New Hampshire, and along a coastal corridor in Maine. Only after 12,000 yr B.P. did the taxa spread northward through the rest of the area. Different tree species entered the southern part of the area at different times and continued their spread at different rates. The approximate order of arrival follows: poplars (13,000–12,000 yr B.P. in the south), spruces, paper birch, and jack pine, followed by balsam fir and larch, and possibly ironwood, ash, and elm, and somewhat later by oak, maple, white pine, and finally hemlock (10,000–9000 yr B.P. in the south).

By mapping and summarizing 478 pollen counts from surface samples at 406 locations in eastern North America, this study documents the relationships between the distributions of pollen and vegetation on a continental scale. The most common pollen types in this region are pine, birch, oak, and spruce. Maps showing isopercentage contours or isopolls for 13 important pollen types reflect the general N-S zonation of the vegetation. The maps and tabulations of average pollen spectra for the six major vegetational regions indicate high values for the following pollen types in each region: (1) tundra-nonarboreal birch, sedge, and alder; (2) forest/tundra-spruce, nonarboreal birch and alder; (3) boreal forest-spruce, jack pine (type), and arboreal birch with fir in the southeastern part; (4) conifer/hardwood forest-white pine, arboreal birch, and hemlock with beech, maple, and oak in the southern part; (5) deciduous forest-oak, pine, hickory, and elm, with beech and maple in the northern part, and highest values of oak and hickory west of the Appalachian crest; and (6) southeastern forest-pine, oak, hickory, tupelo, and Myricaceae. In some cases, less abundant pollen types are diagnostic for the region, e.g., bald cypress in the southeast. In the conifer-hardwood region and southward, pollen of weeds associated with deforestation and agriculture is abundant. The maps also show that much of southeastern U.S. and the area just to the east of Hudson Bay are in need of additional sampling. At 51 of the sites, absolute pollen frequencies (APF; grains/ml lake sediment) were obtained. These confirm the major conclusions from the percentage data, but differences are evident, e.g., the percentages of alder pollen peak in the tundra whereas alder APFs peak in the boreal forest, and spruce percentages peak in the forest-tundra whereas spruce APFs peak in the boreal forest. Because the APF data reflect the patterns of absolute abundance of individual taxa in the vegetation as well as the overall forest densities, future counts of modern pollen should include APF determinations. The effects of sedimentation processes on APF quantities indicate that APF samples should be obtained from moderate size lakes of similar morphology and hydrology and that, in each lake, several samples from the profundal zone should be pooled to create a sample representative of that lake.

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The changing character of vegetation and the effects of disturbance on vegetation are inferred from pollen, plant macrofossils, charcoal, and microlepidopteran larvel head capsules in sediment cores from Upper South Branch Pond, Maine. Following deglaciation 12 500 – 12 000 years ago, a herb–shrub tundra developed which included moss species characteristic of calcareous, mineral soils. Fire and infestation by microlepidopterans were unimportant initially but became important upon arrival of spruce, paper birch, balsam fir, white pine, and tamarack trees (ca. 10 200 – 9500 years BP). Fires were infrequent in the watershed between 7500 and 5000 years BP. The relatively stable forests of this period, dominated by hemlock and yellow birch, grew in what may have been the moistest part of the Holocene. The maximum postglacial abundance of microlepidopteran larvae is centered around the hemlock decline (ca. 4800 years BP). Subsequently, the forest was composed largely of deciduous trees and white pine. Fire incidence was greater, and fewer fossils of microlepidoptera were deposited. Lack of major disturbances between ca. 3300 and 2600 years BP coincided with increases in hemlock, tamarack, yellow birch, and arbor vitae. Increases in boreal conifers began by about 1700 years ago, suggesting cooler, and perhaps wetter, climate. An increase in microlepidoptera accompanied the recent expansion of boreal conifers.

Sediment cores from nine lakes in southern Norway (N) and six in northern New England (NE) were dated by 37 Cs, 210Pb and in NE also by pollen, and were analyzed geochemically and for diatoms. Cores from two N and three NE lakes were analyzed for cladocerans. 37 Cs dating is unreliable in these lakes, probably due to mobility of Cs in the sediment. In Holmvatn sediment, an up-core increase in Fe, starting ca. 1900, correlates with geochemical indications of decreasing mechanical erosion of soils. Diatoms indicate a lake acidification starting in the 1920's. We propose that soil Fe was mobilized and runoff acidified by acidic precipitation and/or by soil acidification resulting from vegetational succession following reduced grazing. Even minor land use changes or disturbances in lake watersheds introduce ambiguity to the sedimentary evidence relating to atmospheric influences. Diatom counts from surface sediments in 36 N and 31 NE lakes were regressed against contemporary water pH to obtain coefficients for computing past pH from subsurface counts. Computed decreases of 0.3-0.8 pH units start between 1890 and 1930 in N lakes already acidic (pH 5.0-5.5) before the decrease. These and lesser decreases in other lakes start decades to over a century after the first sedimentary indications of atmospheric heavy metal pollution. It is proposed that the acidification of precipitation accompanied the metal pollution. The delays in lake acidification may be due to buffering by the lakes and watersheds. The magnitude of acidification and heavy metal loading of the lakes parallels air pollution gradients. Shift in cladoceran remains are contemporary with acidification, preceding elimination of fishes.

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