The Lower Ordovician Fezouata Konservat-Lagerstätte from southern Morocco has been one of the major palaeontological discoveries of the last decade. It provides a unique insight into one of the most critical periods in the evolution of marine life: the Cambrian-Ordovician transition. However, its potential for deciphering key trends in animal diversification was hitherto largely limited by major uncertainties concerning its stratigraphic position, age and environmental setting. Based on extensive fieldwork, fossil evidence, and facies recognition, our study provides clarification on these three crucial issues. Exceptional preservation is limited to two intervals within the Fezouata Shale. Graptolites indicate a late Tremadocian age for the Fezouata Konservat-Lagerstätte as a whole, which is supported by biostratigraphical evidence provided by acritarchs. Sedimentological features and reconstructed patterns of relative sea-level changes indicate relatively shallow-water environmental conditions, under distal storm influence, in an offshore to lower shoreface siliciclastic ramp setting. The Fezouata Biota represents a unique and exceptional window into the palaeobiodiversity in open-marine conditions, thus contrasting with the other Ordovician Konservat-Lagerstätten presently known. In our analyses of this new set of data, we pave the way for accurate temporal, faunal and environmental comparisons with other Lower Palaeozoic Konservat-Lagerstätten, and unlock the full potential of the Fezouata Biota to better understand the processes and scenarios of early animal radiations.

Hybrid depositional systems are created by the interaction of two or more hydrodynamic processes that control facies distribution and their characteristics in terms of sedimentary structures and depositional geometry. The interaction of wave and tide both in the geological sedimentary record and modern environments has been rarely described in the literature. Mixed coastal environments are identified by the evidence of wave and tidal structures and are well identified in nearshore environments, while their recognition in lower shoreface–offshore environments lacks direct information from modern settings. Detailed field analyses of 10 stratigraphic sections of the Lower Ordovician succession (Fezouata and Zini formations; Anti‐Atlas, Morocco) have allowed the definition of 14 facies, all grouped in four facies zones belonging to a storm‐dominated, wave‐dominated sedimentary siliciclastic system characterized by symmetrical ripples of various scales. Peculiar sedimentary organization and sedimentary structures are observed: (i) cyclical changes in size of sedimentary structures under fair‐weather or storm‐weather conditions; (ii) decimetre‐deep erosional surfaces in swaley cross‐stratifications; (iii) deep internal erosion within storm deposits; (iv) discontinuous sandstone layers in most depositional environments, and common deposition of sandstones with a limited lateral extension, interpreted to indicate that deposition at all scales (metric to kilometric) is discontinuous; (v) combined flow–oscillation ripples showing aggrading–prograding internal structures alternating with purely aggrading wave ripples; and (vi) foreshore environments characterized by alternating phases of deposition of parallel stratifications, small‐scale and large‐scale ripples and tens of metres‐wide reactivation surfaces. These characteristics of deposition suggest that wave intensity during storm‐weather or fair‐weather conditions was continuously modulated by another controlling factor of the sedimentation: the tide. However, tidal structures are not recognized, because they were probably not preserved due to dominant action of storms and waves. A model of deposition is provided for this wave‐dominated, tide‐modulated sedimentary system recording proximal offshore to intertidal–foreshore environments, but lacking diagnostic tidal structures.