The development of socio-economic activity over the past ten years in the Mediterranean region has induced severe changes in the main natural forest ecosystems.In the northern Mediterranean, rural depopulation has accelerated since the end of the second World War, particularly since the establishment of Common Market agricultural policies, and led to an under-utilization of species causing a strong biological resurgence of the forest, even at high altitudes. This means that, at the present time, the extension of expansion model coniferous forests is favored by their capacities for spatial, biological and ecological selection. Along with this, the under-utilization of sclerophyllous (resistance model) and deciduous (stabilization model) oak coppices has led to the establishment of new forest structures and architectures which are notably different from the main climatic groups defined up to now by phytosociological and synchronic methods. Two new forms of disturbances have appeared: -increasingly important wild fires have replaced disturbances caused by burn beating and are at the origin of the very strong spatial and temporal heterogeneity of current forest species. -In addition, the geographical continuity of the main groups of same-aged sclerophyllous and deciduous species, due to their non-use over the past ten years, has accelerated a phytosanitary imbalance by an increase in the action of pests.In the southern Mediterranean, particularly in North Africa, demographic pressure and grazing have widely disturbed the main forest ecosystems which show a continual regression of their surface. Many forest tree species with a low spatial and biological selection, such as Mediterranean firs and black pines (Pinus nigra subsp, mauritanica), are threatened with extinction, as are the deciduous oak forests which, considering the climatic stress and edaphic constraints, are permanently in a state of imbalance. Human disturbances induce a complete modification of structures and architectures tending towards the installation of simplified forest models (trees-grasses) where tree regeneration is nearly impossible. The sclerophyllous coppices well-adapted to stress are also threatened by shorter and shorter cutting cycles and by the high usage of tree canopies for grazing. -The forest understory structures have witnessed a decrease in their characteristic sylvatic species and the matorralization of most of the forests can be seen by the replacement of typical forest groups by preforest groups (Tetraclinis forests, Aleppo pine forests). 152-New geopedological constraints linked to the removal of the surface soil layer combined with regular climatic stress (duration of drought periods) strongly decrease the resilience of these ecosystems which are under continual pressure (unbalanced models).-In diverse regions, particulary in semi-arid bioclimates, hyperdegradation affects the shrub cover which disappears for a time in favor of perennial grasses (forest steppization): Andropogon div. sp., Ampelodesmos, Stipa div. sp.In all bio...

Abstract. The natural origin of old Quercus ilex (holm oak) forests on the west coast of Corsica is a matter of dispute. This paper discusses the use of pedoanthracology, especially adapted to approach topics in palaeoecology requiring a high spatial precision. It also shows the importance of fire in vegetation change during the Holocene. Pedoanthracology relies on botanical identification and AMS 14C dating of micro‐charcoal found in soil. Three test pits were dug in a forest currently protected within the ‘Man and Biosphere’ program. These pits yielded large quantities of charcoal. There is evidence that Pinus nigra ssp. laricio (laricio pine) and understorey heather species (Ericaceae) played an important role in the vegetation at the beginning of the Subatlantic (ca. 2500 BP). The importance of Pinus pinaster (maritime pine), Quercus ilex and a few mesophilic species increased between 2000 and 1000 BP. Charcoal fragments found in the soil show the role of fire in the vegetation changes recorded. The late expansion of P. pinaster results from a high fire frequency. The hypothesis proposed in the paper suggests that fires destroyed the remaining toxic humus that had accumulated during the millennia which preceded the arrival of Q. ilex to our study sites. The late development of holm oak forest is a combination of the migration of the species and the increase of fire frequency since 2500 BP, most probably resulting from slash‐and‐burn agriculture.

Quercus ilex sensu lato plays an important role in Western Mediterranean ecosystems, but is poorly developed in the Eastern Mediterranean where it is often replaced by Quercus calliprinos.The occurrence of Quercus ilex in the different bioclimates and their thermic subdivisions is presented on a small scale. Under certain geographical and ecological conditions, Quercus ilex participates in the organization of vegetation ecosystems from the meso-Mediterranean to the oro-Mediterranean altitudinal zones.Quercus ilex imposes microclimatic constraints on its associated species; it thus organizes an original understory vegetation structure (ethoh)gical groups) which will be defined in this study.One of the reasons for the success of Quercus ilex stems from its remarkable resistance to ecological constraints. A broad synthetic presentation of foliar area index variations in relation to different types of stress serve as a basis for an explanation of the sclerophyllous model in the Mediterranean region.Finally, historical factors are outlined as being critical characteristics in the determination of the present organization and spatial structure of Quercus ilex ecosystems. These considerations will be analyzed from a paleobioclimatical point of view, including data related to human pressure. Distribution and biogeography of sclerophyllous oaks (Fig.

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