The phylogeographical evolution and the consequent changing distribution and diversity of rhynchonelliform brachiopods through the Ordovician are linked to the dynamic palaeogeography of the period. The Early Ordovician (Tremadocian and Floian) is characterized by globally low-diversity faunas with local biodiversity epicentres, notably on the South China Palaeoplate; low-latitude porambonitoid-dominated faunas with early plectambonitoid and clitambonitoid representatives, as well as high-latitude assemblages mostly dominated by orthoids, can be recognized, but many taxa are rooted in Late Cambrian stocks. The Early Ordovician displays a steady increase in rhynchonelliformean biodiversity, which was mostly driven by the increasing success of the Porambonitoidea and Orthoidea, but the billingsellids and early plectambonitoids also contributed to this expansion. During the Early to Mid Ordovician (Dapingian–Darriwilian), marine life experienced an unprecedented hike in diversity at the species, genus and family levels that firmly installed the suspension-feeding benthos as the main component of the Palaeozoic fauna. However, this may have occurred in response to an early Darriwilian annihilation of existing clades, some of which had been most successful during the Early Ordovician. New clades rapidly expanded. The continents were widely dispersed together with a large number of microcontinents and volcanic arcs related to intense magmatic and tectonic activity. Climates were warm and sea-levels were high. Pivotal to the entire diversification is the role of gamma (inter-provincial) diversity and by implication the spread of the continents and frequency of island arcs and microcontinents. The phylogeographical analysis demonstrates that this new palaeogeographical configuration was particularly well explored and utilized by the strophomenides, especially the Plectambonitoidea, which radiated rapidly during this interval. The porambonitoids, on the other hand, were still in recovery following the early Darriwilian extinctions. Orthides remained dominant, particularly at high latitudes. Biodiversity epicentres were located on most of the larger palaeoplates, as well as within the Iapetus Ocean. Provincial patterns were disrupted during the Sandbian and early Katian with the migration of many elements of the benthos into deeper-water regimes, enjoying a more cosmopolitan distribution. Later Katian faunas exhibit a partition between carbonate and clastic environments. During the latest Katian, biogeographical patterns were disrupted by polewards migrations of warm-water taxa in response to the changing climate; possibly as a consequence of low-latitude cradles being developed in, for instance, carbonate reef settings. Many clades were well established with especially the strophomenides beginning to outnumber the previously successful orthides, although this process had already begun, regionally, in the mid to late Darriwilian. At the same time, atrypoid and pentameroid clades also began to radiate in low-latitude faunas, anticipating their dominance in Silurian faunas. The Hirnantian was marked by severe extinctions particularly across orthide-strophomenide clades within the context of few, but well-defined, climatically controlled provincial belts.Supplementary material:The individual localities and a reference list for the data sources are provided at:http://www.geolsoc.org.uk/SUP18667

Analysis of the Upper Jurassic NaknekFormation in the Talkeetna Mountains, Alaska, documents synorogenic sedimentation in a forearc basin along the outboard (southern) margin of the allochthonous Peninsular terrane during accretion to the western North American continental margin. New geochronologic, sedimentologic, and compositional data defi ne a two-part stratigraphy for the Naknek Formation. Microfossil, megafossil, and U-Pb clast ages document early Oxfordian to early Kimmeridgian deposition of the lower 690 m of the Naknek Formation and early Kimmeridgian to early Tithonian deposition of the upper 225 m of the Naknek Formation. Lithofacies and paleocurrent data from the lower Naknek Formation demonstrate initial deposition on a high-gradient, southward-dipping basin fl oor. Submarine mass fl ows deposited poorly sorted, cobble-boulder conglomerate in proximal fan-delta environments. Gravelly mass fl ows transformed downslope into sandy turbidity currents on a muddy prodelta slope. During early Kimmeridgian to early Tithonian time, fan-delta environments were replaced by lower gradient marine shelf environments characterized by deposition of cross-stratifi ed sandstone and bioturbated mudstone. Source-diagnostic clasts, feldspathic sandstone compositions, southwarddirected paleocurrent indicators, and U-Pb zircon ages of plutonic clasts (167.6 ± 0.3 Ma; 166.5 ± 0.2 Ma, 164-159 Ma, 156.2 ± 0.4 Ma)indicate that the Naknek Formation was derived primarily from volcanic and plutonic source terranes exposed along the northern basin margin in the southern Talkeetna Mountains. Geologic mapping documents the Little Oshetna fault, a newly identifi ed northward-dipping reverse fault that bounds the northern margin of the Naknek Formation in the Talkeetna Mountains. The concentration of boulder-rich mass-fl ow deposits in the footwall of the fault in combination with geochronologic and compositional data suggest that sedimentation was coeval with Late Jurassic shortening along the fault and exhumation of plutonic source terranes exposed in the hanging wall of the fault. From a regional perspective, coarse-grained forearc sedimentation and pluton exhumation along the outboard (southern) segment of the Peninsular terrane were coeval with crustal-scale shortening and synorogenic sedimentation in retroarc basins along the inboard (northern) margin of the Wrangellia terrane (Kahiltna, Nutzotin, and Wrangell Mountains basins). We interpret the regional and synchronous nature of Late Jurassic crustal-scale deformation and synorogenic sedimentation in south-central Alaska as refl ecting either initial collision of the Wrangellia and Peninsular terranes with the former continental margin of western North America or amalgamation of the two terranes prior to collision.

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Mesozoic strata of the northwestern Talkeetna Mountains are located in a regional suture zone between the allochthonous Wrangellia composite terrane and the former Mesozoic continental margin of North America (i.e., the Yukon-Tanana terrane). New geologic mapping, measured stratigraphic sections, and provenance data define a distinct three-part stratigraphy for these strata. The lowermost unit is greater than 290 m thick and consists of Upper Triassic-Lower Jurassic mafic lavas, fossiliferous limestone, and a volcaniclastic unit that collectively we informally refer to as the Honolulu Pass formation. The uppermost 75 m of the Honolulu Pass formation represent a condensed stratigraphic interval that records limited sedimentation over a period of up to ca. 25 m.y. during Early Jurassic time. The contact between the Honolulu Pass formation and the overlying Upper Jurassic-Lower Cretaceous clastic marine strata of the Kahiltna assemblage represents a ca. 20 m.y. depositional hiatus that spans the Middle Jurassic and part of Late Jurassic time. The Kahiltna assemblage may to be up to 3000 m thick and contains detrital zircons that have a robust U-Pb peak probability age of 119.2 Ma (i.e., minimum crystallization age/maximum depositional age). These data suggest that the upper age of the Kahiltna assemblage may be a minimum of 10-15 m.y. younger than the previously reported upper

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