This paper examines the mid-to late-Holocene (c. 6850-1160 cal. yr BP) environmental history of Sierra de Gádor in southern Spain. The local vegetation dynamics are reconstructed through the palaeoecological record obtained from a lacustrine deposit situated at 1530 m. Various hypotheses are considered to explain the vegetation dynamics apparent in the palaeoecological evidence, including climatic change, re occurrence and human activity. Although the vegetation in this region is sensitive to climatic change, threshold events driven by ecological factors are also apparent. Climatic events include a thermo-mesophytic optimum with abundance of deciduous trees and maximum lake water level found from c. 6850 to 5500 cal. yr BP. In contrast, changes in the frequency of major re episodes appear to have shaped interspeci c relationships and vegetation change, especially from c. 4200 cal. yr BP onwards. Biotic properties of the ecosystem such as the inertia of established tree populations, interconnected with competition adjustments, appear also to have played a role. Over the last two millennia, overgrazing, combined with natural and/or human-set res, appears to have pushed mountain forests over a threshold leading to the spread of grassland, thorny scrub, junipers and nitrophilous communities.

Sima del Elefante, Atapuerca, Spain contains one of the earliest hominin fragments yet known in Europe, dating to 1.2 Ma. Dental calculus from a hominin molar was removed, degraded and analysed to recover entrapped remains. Evidence for plant use at this time is very limited and this study has revealed the earliest direct evidence for foods consumed in the genus Homo. This comprises starchy carbohydrates from two plants, including a species of grass from the Triticeae or Bromideae tribe, meat and plant fibres. All food was eaten raw, and there is no evidence for processing of the starch granules which are intact and undamaged. Additional biographical detail includes fragments of non-edible wood found adjacent to an interproximal groove suggesting oral hygiene activities, while plant fibres may be linked to raw material processing. Environmental evidence comprises spores, insect fragments and conifer pollen grains which are consistent with a forested environment.

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