We used a Simrad EK60 echosounder equipped with two split-beam transducers to develop a sampling strategy for assessing fish resources in Tunisian man-made lakes. Day and night surveys, using vertical and horizontal beaming, were carried out between December 2006 and February 2007, a period when fish catchability is high. Four reservoirs with differing surface areas and bathymetries were selected. Echogram analysis revealed that fish communities were mainly composed of individual targets. A few schools were detected near the surface during daylight, but these schools dispersed slightly at night. In these multispecies reservoirs, considerable day and night differences in density existed, but with no clear trend. Target strength (TS) distribution mode values detected at night were always lower or equal to daytime values. Biomass estimates were significantly higher during daytime in three reservoirs, corresponding with higher TS modal values. In the other reservoir, the biomass estimate was significantly higher during nighttime corresponding with higher mean density during this period. Using only a vertically aimed transducer in our study reservoirs would have led to an underestimate of density and biomass by 5-100% and 20-100%, respectively, depending on the man-made lake. We conclude that acoustic sampling in our reservoirs must be done during day and night and that both vertical and horizontal beaming must be used to obtain the best possible picture of the fish stocks.
In order to test the hypothesis that fish distribution is related to water depth, three exploited reservoirs were sampled at day and at night using a Simrad EK60 echosounder. Acoustic surveys were carried out between February and June 2008, which focused on the fish size, species density (fish per ha) and biomass (kg ha )1 ) along vertical and longitudinal gradients. It was evident in the surface layer (0-3 m) during daytime that the larger fish sizes occupy waters near the dam or the middle of Lakhmess and Sejnane reservoirs. In the upper layer at nighttime, a gradient of fish size is shown proportional to the depth at Lakhmess Reservoir and inversely proportional at the man-made Sejnane Lake. At Lakhmess and Laabid reservoirs, fish density was significantly higher in deep water (3926 ± 1409 and 624 ± 258 fish per ha) rather than in the upper layer (988 ± 2 and 8 ± 2 fish per ha in daytime, respectively), while at Sejnane Reservoir the number of fish per area was similar along the vertical gradient. The biomass was significantly higher in waters deeper than 3 m at Sejnane Reservoir whereas at Lakhmess it was higher in the surface layer. Fish biomass increased from upstream (0.16 ± 0.05 kg ha )1 ) to middle (3.07 ± 2.96 kg ha )1 ) and downstream (3.82 ± 2.30 kg ha )1 at night) areas in the man-made Laabid Lake while a similar longitudinal gradient occurred in the deepest Sejnane Reservoir.
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