The nuclear ITS1-5.8S-ITS2 DNA region is studied for 29 specimens of Coscinodon from Eurasia, mostly from Russia, and 4 specimens from North America. The sequences were found to be much less variable than in two other genera of the Grimmiaceae, Grimmia and Schistidium. Coscinodon yukonensis Hastings was found to be rather widespread in Asia, despite never having been reported before in this continent, being erroneously treated as C. humilis Milde. European C. humilis is quite distinct molecularly from Asian C. yukonensis and forms a sister clade to C. cribrosus (Hedw.) Spruce. Some Asian populations of "C. cribrosus" were unresolved, being found outside the main clade formed by C. cribrosus and C. humilis. Coscinodon hartzii C.E.O. Jensen forms a basal grade to the rest of the species in the genus. Coscinodon pseudohartzii is described as a species new to science from Siberia. РезюмеИзучен участок ITS1-5.8S-ITS2 ядерной ДНК у 29 образцов рода Coscinodon из Евразии, б.ч. из России, и 4 образцов из Северной Америки. Последователь-ности оказались намного менее вариабельными по сравнению с таковыми у двух других родов Grimmiaceae, Grimmia и Schistidium. Coscinodon yukonensis Hastings ранее не был отмечен в Азии, ранее образцы этого вида с этого континента относили к C. humilis Milde. Типичный C. humilis из Европы, однако, молекулярно отличается от него очень сильно и в то же время более близок к C. cribrosus (Hedw.) Spruce. Европейские и часть азиатских популяций C. cribrosus образуют кладу, сестринскую C. humilis, но ряд азиатских популяций "C. cribrosus" оказывается в неразрешенном положении. Coscinodon hartzii C.E.O. Jensen образует граду в основании дерева рода. По результатам анализа в Сибири выявлен не описанный ранее вид, C. pseudohartzii.
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