The Eocene-Oligocene (E-O) transition was an epoch of great turnover, in both terrestrial and marine environments. Mammals were highly affected by an extinction event called grande coupure of Stehlin, which has been very well documented in Europe and Asia (Hartenberger, 1998). In America the tropical molluscs underwent a significant extinction event at the beginning of the Oi-1 glaciation in the Oligocene (Hickman, 2003). This drop in temperature was caused by the opening of the Drake Strait, thereby giving rise to the circum Antarctic current, the formation of ice caps on the poles and the development of the psychrosphere in
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