With 1 plate and 12 figures in the text) Newts were collected throughout the year from both breeding ponds and terrestrial sites and were weighed, measured and dissected; in males, the testes were examined histologically. Smooth newts show post-nuptial gametogenesis such that, during late summer and autumn they are producing mature gametes for the following year's breeding season. In males, the testes are at their smallest size during the spring, when they consist mostly of immature sperm cysts and evacuated tissue, mature sperm having been evacuated into the vasa deferentia during the newts' migration to water. Evacuated testicular tissue is glandular in function and there is evidence that secretions from this tissue control the development of secondary sexual characters: the dorsal crest, fringes of skin on the toes and the dorsal cloaca1 gland. In both sexes, fat body and liver weights are lowest in the spring and increase in the autumn. In females, oocytes vary in size, depending on the amount of yolk they contain. Only the larger oocytes are laid in a current breeding season, the smaller ones being retained and yolked in late summer and autumn. In both sexes, measures of fecundity (testis size in males, oocyte number in females) are strongly correlated with body size. The finding that male newts have a finite supply of sperm during the breeding season leads to an interpretation of various aspects of male courtship behaviour. These are adaptations for conserving sperm and allocating it to courtship encounters in a way likely to promote male reproductive success.

In relation to the size of its population,Wales has a relatively large number of higher education institutions, several of which are small and located in rural areas. Compared with other parts of the United Kingdom, only a small number of higher education students are taught in Welsh further education colleges. Nevertheless, efforts have been made to increase higher education provision in the colleges, beginning in the late 1980s with the encouragement of franchising arrangements by the Wales Advisory Body and later with a joint initiative by the Higher and Further Education Funding Councils for Wales to expand some sub-degree higher education in the colleges by direct funding. These developments represent just one strand of activity linking the higher and further education sectors in Wales and have not been a subject of major or regular policy attention. For reasons of scale and geography, and through the enabling structures and processes serving both funding councils, cross-sector initiatives and collaboration have generally been given high priority in strategies to widen access and build progression. The administrative devolution reflected in these arrangements has been increased by political devolution since 1999, with as yet unclear implications for the future development of higher education in the colleges.