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S elatior in Britain1," I had the pleasure of showing to Dr Bateson, Miss E. R. Saunders, and others the various Primula hybrids I had described therein-namely : (I) P. elatior x vulgaris, ( 2 ) P. elatior x veris, and (3) P. veris x vulgaris-all growing in a wild state in certain woods in Essex, Suffolk, and Cambridgeshire.On this occasion Dr Bateson (without disputing my assumption that the plants in question really were hybrids) pointed out that, for purposes of absolute scientific accuracy, their hybrid origin, even if practically certain inferentially, ought to be demonstrated incontestibly by means of definite experiment.Thereupon it was decided to attempt to accomplish this by planting some Oxlips (P. elatior) in woods wherein Primroses (P. vulgaris) only were growing and some Primroses in other woods wherein Oxlips only were growing, and then waiting to see whether plants similar to those I had described as hybrids between these two species would appear naturally in the course of time.That the plants I had described as hybrids were really such might, of course, have been demonstrated much more easily and quickly by artificially pollinating flowers of either of the two reputed parents with pollen from flowers of the other on covered plants growing in cultivation and then raising plants from any fertile seed resulting. As a matter of fact, Miss Saunders succeeded in doing this soon after the time when our experiments described herein were begun2, and since then Mr E. G. Highfield has done the * Miss Saunders writes me: '' In 1899 and rgoo.1 raised a number of hybrids between P. elatior and P. vulgaris from crosses which I made for the purpose of testing the views expressed in your original paper. The results entirely confirmed your conclusions; but, having done this and having found the sterility so great that there was no likelihood of obtaining any genuine ratios in support of the Mendelian Theory or otherwise, I did not continue the experiments and no record was published." Journ. Linn.