Constraining effects of seasonal cold are obvious in temperate zone habitats, and effects of seasonal drought can be obvious in areas of highly seasonal rainfall. But phenological patterns become more variable, subtle, and complex in less seasonal tropical environments (Abernethy et al., 2018). Many studies of phenology emphasize plants, but phenology of consumers also receives attention (Van Schaik et al., 1993). Tropical insects exhibit a wide range of seasonal patterns, even in very weakly seasonal regions (Kishimoto-Yamada et al., 2015;Wolda, 1988). Ants are dominant consumers in tropical forests and phenology of ant reproductive flights has been
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