Between 2010 and 2011 a field survey dedicated to Dracaena cinnabari (DC) population was conducted in Firmihin, Socotra Island (Yemen). It’s main goal was to collect data that would make it possible to unbiasedly estimate main characteristics of the local DC population. Our motivation was to provide reliable information to support decision-making processes as well as other research activities. At the same time we were not aware of a survey which could provide this kind of statistical-sound estimates for the whole population covering an area of almost 700 ha.
This article describes how the survey has been planned and carried out in practice. In addition, we also provide a set of preliminary estimates of the main DC population figures - totals and per hectare densities of stems, overall and partitioned according to predicted crown age. Among estimated parameters there are also mean crown age and proportions of predefined age classes on the total number of living DC stems. These estimates provide an explicit information on age structure of the whole DC population in Firmihin.
Although we collected data on more than one hundred randomly located plots, the reported accuracy of our estimates is still rather limiting. We discuss several possibilities to obtain more accurate results or at least to approach the supposedly lower true variance that can’t be calculated by approximate techniques applied here.
The design and concept of our survey makes it possible to evaluate changes over time on stem by stem bases and to generalize these stem-level details to the whole population. Mortality, regeneration and even change of population’s mean crown age can be estimated from a future repeated survey, which would be extremely useful to draw firm conclusions about the dynamic of the whole DC population in Firmihin.
Questions
What was the main trigger of treeline ecotone advance – rising temperature or agricultural land abandonment? Were the triggering factors of tree expansion homogeneous or did they differ between upper and lower parts of the treeline ecotone?
Location
Sudetes Mts., Central Europe (50° N, 15‐17° E).
Methods
Data were gathered from the treeline ecotone formed by Picea abies at elevations ranging from 1250 to 1490 m. The study area experienced a 1 °C temperature increase over the last 100 yr and termination of cattle grazing and grass mowing in the first half of the 20th century. At 38 plots situated at lower (‘timberline’) and middle or upper (‘treeline’ and ‘outpost treeline’) parts of the treeline ecotone, the age structure of all seed origin P. abies was determined. Changes in tree cover and number of trees over the last 60–70 yr were assessed from aerial imagery. The history of agricultural land use for each plot was compiled. Finally, changes in tree establishment were modelled using climatic variables and land‐use intensity.
Results
We found that tree establishment at treeline had occurred with a 30–40‐yr lag after the main establishment peak at timberline. Whereas all treeline plots showed gradual increases in tree cover, timberline tree cover first increased, with some plots then undergoing thinning. Enhanced tree establishment was dependent mainly upon agricultural land abandonment. The effect of land‐use changes was more important in the lower than in the upper part of the treeline ecotone. Increasing summer temperatures had a negative influence on seedling establishment in the last few decades.
Conclusions
Treeline ecotone densification was attributable to agricultural land abandonment across the entire treeline ecotone with the most important effect at timberline. More recently, seedling establishment has been limited by the effects of drought and/or absence of suitable microsites. We documented that one or two establishment pulses over 120 yr, together with enhanced growth since the 1980s, were able to trigger treeline ecotone advance.
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