Summary 1.In some ecosystems there is a positive feedback between forage quality and grazing intensity. This involves three components of plant tolerance to grazing: functional traits, herbivore selectivity and response to grazing. We analysed the relationships between these components at species and community levels in Patagonian steppe grasslands. 2. We measured plant functional traits [height, specific leaf area (SLA) and foliar toughness] and estimated sheep selectivity and grazing response indices for 35 plant species. Sheep selectivity indices were obtained from microhistological and species' availability data, and grazing response indices from species' abundances in sites with contrasting grazing intensities. We performed correlations and multiple regressions among the three types of variables across the pool of 35 species. 3. To analyse data at the community level, we computed weighted averages of traits and sheep selectivity indices for 34 floristic samples taken from each side of 17 fence lines with contrasting grazing intensities. Correlations between mean trait values and sheep selectivity across the 34 samples, and paired comparisons of those variables between sides of the fences, were performed. 4. Taller plants had leaves with lower SLA and/or higher toughness. Short species of intermediate toughness were selected more often by sheep, while SLA was not related to sheep selectivity. Short species with high SLA increased with grazing, while toughness and sheep selectivity were unrelated to grazing response. 5. At the community level, short swards with high average SLA had high selectivity indices and were more abundant on the most intensively grazed sides of fence lines. Leaf toughness was unrelated to other traits or to sheep selectivity, and showed no significant response to grazing. 6. Synthesis and applications . Intensive grazing can increase the forage value of grasslands by the creation of lawns dominated by tolerant species. However, results from this study showed that some plant species that were avoided by grazers also increased, indicating a potential risk of a shift in composition of grazing lawns towards states of low forage value. This suggests that periodic resting of lawns could be a good management strategy to favour palatable species, thereby minimizing the risk of undesirable shifts in the overall species composition.

Sheep diets were determined seasonally for large flocks grazing year-round in 2 landscape types of the Magellanic steppe of Argentina. A tussock-grass steppe of Festuca gracillima Hooker f. dominates the uplands of the whole area. On acid soils (Quaternary landscape), woody variants of the steppe prevail; on neutral soils (Tertiary landscape), woody plants are almost ahsent and short grasses and forbs are abundant. Principal tasa consumed throughout the year were: Poa L., Deschampsia P.Beauv., and "sedges & rushes". Consumption of woody species and of the dominant tussock-grass Festuca gracillima increased notably in winter. Despite the large proportion of species in common, diets differed significantly between landscapes. In the Quaternary landscape, which has a higher botanical diversity, diets were more dissimilar among seasons and had a higher annual diversity indes. Because of their different composition of forage types the 2 landscapes differed in their overall grazing value. The Tertiary landscape, with a low floristic diversity but richer in highly preferred species as Poa spp. would be a more risky grazing area in winter, when an ice sheet or a snow cover limits harvesting of the lower layer of short grasses and forbs.

Abstract. In the Festuca gracillima tussock steppe of northern Tierra del Fuego there is a physiognomic pattern at the community scale composed of a matrix of tussocks, scrub patches of Chiliotrichum diffusum and lawn patches. In this paper we compared floristic composition, soils, microclimate and sheep use of these three types. Species composition greatly differed among physiognomic types. Lawn patches showed the highest diversity and maximum cover of exotics, grazing escape growth forms and species of high‐fertility habitats. Soil profiles of tussock steppe and lawn were similar but the A‐horizon of the latter contained more clay and cations. Radiation and temperature during daylight at the soil surface in summer were highest in the lawn patches. Soil impedance, a good indicator of trampling, was highest on the lawn patches. According to dung pellet density, lawn patches received much heavier use than the tussock matrix. Our results suggest that lawn patches supported the highest grazing impact and undergo a eutrophication process favoured by high mineral inputs, high mineralization rates in the soil during the warm season and restricted leaching through the profile. They also support the idea that a feedback mechanism between vegetation heterogeneity and sheep grazing behaviour maintains the patches. Although grazing behaviour suggests that the lawn patches favour animal production, their expansion would result in a system less protected against the impact of water deficits and temperature variations and more vulnerable to erosion and to the loss of important species.

The cover of Festuca gracillima (coiró n fueguino), a native tussock grass that dominates grass steppes of Southern Patagonia and Tierra del Fuego, has diminished under continuous sheep grazing. This loss is a concern, because it also reduces forage availability in winter, biodiversity, and soil stability. In the present study, the hypothesis that tussock grass birth and mortality rates are balanced only under moderate-grazing or exclusion regimes was tested with two 5-year records of demographic data obtained from 3 sheep grazing regimes: exclosure (no grazing), low (0.0348 AU ha À1 y À1 , where AU represents animal units equivalent to the consumption of a 450-kg cow), and high (0.1043 AU ha À1 y À1). Tussocks were outlined in photographs and marked in the field at two 5-year intervals. The initial total number of plants for the 3 grazing regimes (n ¼ 358) increased to 384 plants at the end of the 10-year period. No recruitment from seed was observed; plant number changed as a balance of yearly rates of tussock mortality (1.48%), amalgamation (0.75%), and subdivision (2.04%). Intensely grazed populations showed greater (2.13%) mortality rates than ungrazed (1.20%) or moderately grazed (0.78%) populations. Tussocks in intensely grazed populations were smaller (167 cm 2 plant À1) than those in moderately grazed (197 cm 2 plant À1) or ungrazed (300 cm 2 plant À1) populations. Transition matrices showed eigenvalues of 0.701 (high grazing), 0.794 (exclosure), and 0.876 (low grazing). All growth rates of demographic models were negative; the largest rate of population decrease was found under high-intensity grazing, for which projections show that half the tussocks would be lost in 37 years. Under low-intensity grazing and exclosure, a similar tussock loss would take place in 87 and 74 years, respectively. Results show 1) the importance of vegetative processes for tussock demography, 2) the extremely slow dynamics of population changes, and 3) that tussocks may suffer increased mortality as a consequence of subdivision or fragmentation, a process that can be viewed as a small-scale example of the generalized effect of patch subdivision under grazing. Resumen Festuca gracillima (coiró n fueguino) es una gramínea mediana dominante en los pastizales del sur de la Patagonia y norte de Tierra del Fuego que ha ido perdiendo cobertura bajo pastoreo continuo. El proceso implica también una reducció n en el forraje disponible en invierno, en la estabilidad del suelo y en la biodiversidad. El presente trabajo analiza 10 añ os de demografía de poblaciones de coirones sujetos a pastoreo controlado con tres intensidades: clausura (sin pastoreo), alta (0.1043 AU ha À1 añ o À1 , AU son unidades animales equivalentes al consumo de una vaca de 450 kg), y baja (0.0348 AU ha À1 añ o À1). Se puso a prueba la hipó tesis de que las tasas de natalidad y mortalidad llegan al equilibrio solamente bajo pastoreo moderado o en exclusió n. Los coirones fueron identificados en mapas fotográficos y marcados en el campo en dos períodos consecutivos ...

Biological invasions have important ecological impacts at both local and global levels, affecting biodiversity, ecosystem functioning, and economic sustainability. The study of invasions requires specific methodological approaches to gain rapid insight into the spatial and temporal dynamics of the system. We studied a recent invasion by the exotic herb Hieracium pilosella L. (Mouse-ear hawkweed, Asteraceae) in the Argentinean portion of the northern part of Tierra del Fuego Island in Southern Patagonia. To assess the extent of this invasion and the related ecological and land use factors, we performed an extensive field sampling at a regional scale and used a spatial pattern approach using geostatistical techniques to build a map of the invasion. Our results showed that the invading species is widely distributed across the entire Fuegian steppe, in general with low cover (\2%) and a particular spatial structure having hot spots (10-70%) related to specific land uses. We found regionallatitudinal and also local-community level variability in the frequency, cover and spatial distribution of the invasive species, each of them associated with particular ecological factors, and no association with regular domestic animal grazing. Region-wide invasion may be accounted for by wind dispersal of the seeds and the latitudinal variability of the precipitation; while localvariability was associated with the susceptibility of different plant communities to invasion. Hot spots were found to be related to massive soil disturbances (e.g., road building, shrub removal, physical impacts of long-term and intense domestic animal use, etc.), probably due to the presence of bare soil favoring colonization by the invading species. Scrublands, lawns and grasslands were more invaded than wetlands, heathlands or salt grasslands. Since the invasion is apparently at initial stages due to the low cover values found, we believe that information about the extent and environmental or management factors involved in this exotic plant's expansion should be used to design appropriate control and mitigation tools. In this sense, our results point to management actions focused on reducing the presence of hot spots of invasion by means of quick revegetation of bare soil after a massive disturbance event takes place.