The human settlement of Europe during Pleistocene times was sporadic and several stages have been recognized, both from paleaoanthropological and archaeological records. If the first phase of hominin occupation (as early as 1.4 Ma) seems mainly restricted to the southern part of the continent, the second phase, characterized by specific lithic tools (handaxes), is linked to Acheulean settlements and to the emergence of Homo heidelbergensis, the ancestor of Neanderthals. This phase reached northwestern Europe and is documented in numerous sites in Germany, Great Britain and northern France, generally after 600 ka.At la Noira (Brinay, Central France), the Middle Pleistocene alluvial formation of the Cher River covers an archaeological level associated with a slope deposit (diamicton). The lithic assemblage from this level includes Large Cutting Tools (LCTs), flakes and cores, associated with numerous millstone slabs. The lithic series is classified as Acheulean on the basis of both technological and typological analyses. Cryoturbation features indicate that the slope deposits and associated archaeological level were strongly frozen and disturbed after hominin occupation and before fluvial deposition. Eight sediment samples were dated by the electron spin resonance (ESR) method and the weighted average age obtained for the fluvial sands overlying the slope deposits is 665±55 ka. This age is older than previous chronological data placing the first European Acheulean assemblages north of 45th parallel north at around 500 ka and modifies our current vision of the initial peopling of northern Europe. Acheulean settlements are older than previously assumed and the oldest evidences are not only located in southern Europe. La Noira is the oldest evidence of Acheulean presence in north-western Europe and attests to the possibility of pioneering phases of Acheulean settlement which would have taken place on a Mode 1-type substratum as early as 700 ka. The lithic assemblage from la Noira thus provides behavioral and technological data on early Acheulean occupation in Europe and contributes to our understanding of the diffusion of this tradition.

Neanderthals are most often portrayed as big game hunters who derived the vast majority of their diet from large terrestrial herbivores while birds, fish and plants are seen as relatively unimportant or beyond the capabilities of Neanderthals. Although evidence for exploitation of other resources (small mammals, birds, fish, shellfish, and plants) has been found at certain Neanderthal sites, these are typically dismissed as unusual exceptions. The general view suggests that Neanderthal diet may broaden with time, but that this only occurs sometime after 50,000 years ago. We present evidence, in the form of lithic residue and use-wear analyses, for an example of a broad-based subsistence for Neanderthals at the site of Payre, Ardèche, France (beginning of MIS 5/end of MIS 6 to beginning of MIS 7/end of MIS 8; approximately 125–250,000 years ago). In addition to large terrestrial herbivores, Neanderthals at Payre also exploited starchy plants, birds, and fish. These results demonstrate a varied subsistence already in place with early Neanderthals and suggest that our ideas of Neanderthal subsistence are biased by our dependence on the zooarchaeological record and a deep-seated intellectual emphasis on big game hunting.

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Notarchirico (Southern Italy) has yielded the earliest evidence of Acheulean settlement in Italy and four older occupation levels have recently been unearthed, including one with bifaces, extending the roots of the Acheulean in Italy even further back in time. New 40 Ar/ 39 Ar on tephras and ESR dates on bleached quartz securely and accurately place these occupations between 695 and 670 ka (MIS 17), penecontemporaneous with the Moulin-Quignon and la Noira sites (France). These new data demonstrate a very rapid expansion of shared traditions over Western Europe during a period of highly variable climatic conditions, including interglacial and glacial episodes, between 670 and 650 (i.e., MIS17/MIS16 transition). The diversity of tools and activities observed in these three sites shows that Western Europe was populated by adaptable hominins during this time. These conclusions question the existence of refuge areas during intense glacial stages and raise questions concerning understudied migration pathways, such as the Sicilian route. Recent data in prehistory tend to break down the previously supposed clear boundary between anatomically "modern" humans (Homo sapiens), associated with modern cognitive abilities, and earlier hominin species, such as Neanderthals and their ancestors, Homo heidelbergensis. The earliest Acheulean Homo heidelbergensis groups arrived in Europe ca. 1.0-0.7 Ma ago, and were exposed to challenging environmental conditions, which may have stimulated new cultural responses. These included behavioral innovations that could have potentially derived from African or Levantine origins during these early time ranges. During this period, named the Mid-Pleistocene revolution, Northern Europe was characterized by constraining climatic conditions, especially during glacial stages (1.25-0.7 Ma). Major topographic and sea-crossing barriers constituted considerable challenges

Monospecific exploitation of reindeer by Neanderthals is common behaviour in the Upper Pleistocene of WesternEurope. However, reindeer-dominated assemblages have largely been reported from regions of northern Germany and south-western France, with few examples noted in south-eastern France, where faunal assemblages are most of the time dominated by a variety of other large ungulates such as red deer, horse and diverse bovids. Here, we present multi- ManuscriptClick here to download Manuscript Daujeard_et_al_Maras_4.1_Text_AAS.docx strand (bio-and eco-) archaeological datasets from the site of Abri du Maras (level 4.1), situated at the mouth of the Ardèche and Rhône rivers, a new example of a reindeer-dominated Neanderthal site in south-eastern France. Dated to the beginning of the MIS 3, the zooarchaeological assemblage is dominated by reindeer (88% of the NISP, representing 16 individuals), but also includes horse, bison, megaceros, red deer, ibex and lagomorphs. The combination of zooarchaeological, cementochronological and tooth microwear analyses evidence a single-species dominated spectrum, with catastrophic mortality and repeated autumnal deaths. This integrated approach provides an extensive picture of human subsistence behaviour, pointing to short-term hunting episodes of reindeer herds in an exceptional context of a quasi-exclusive Neanderthal accumulation. The high number of individuals and selective butchery may correspond with a cooperative and planned mass hunting strategy. The multidisciplinary approach undertaken here also incorporating paleontological, charcoal, ecological, and isotopic analyses places the archaeological and zooarchaeological data within a broader regional palaeoenvironmental framework, providing valuable landscape-contextual information. The zooarchaeological data suggest a subsistence behaviour different from other Neanderthal reindeer-dominated assemblages often connected with specialised butchery or hunting sites.

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