The fossil record of the earliest tetrapods (vertebrates with limbs rather than paired fins) consists of body fossils and trackways. The earliest body fossils of tetrapods date to the Late Devonian period (late Frasnian stage) and are preceded by transitional elpistostegids such as Panderichthys and Tiktaalik that still have paired fins. Claims of tetrapod trackways predating these body fossils have remained controversial with regard to both age and the identity of the track makers. Here we present well-preserved and securely dated tetrapod tracks from Polish marine tidal flat sediments of early Middle Devonian (Eifelian stage) age that are approximately 18 million years older than the earliest tetrapod body fossils and 10 million years earlier than the oldest elpistostegids. They force a radical reassessment of the timing, ecology and environmental setting of the fish-tetrapod transition, as well as the completeness of the body fossil record.

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The first deep reflection seismic profile was acquired across the Teisseyre‐Tornquist Zone (TTZ), a major European suture zone, southeast of the Avalonia‐Baltica collision zone, portraying different tectonic deformation styles (thick‐ versus thin‐skinned) and Phanerozoic crustal accretion episodes over a relatively short distance (240 km). The overall crustal structure preserves image of the Neoproterozoic passive margin setting but with a significant overprint of the repetitive tectonic convergence (Cadomian to Alpine) acting at the Baltica margin. In contrast to the Avalonia‐Baltica collision zone, here the polygenetic crustal accretion occurring at the cratonic margin was ultimately shaped by significant strike‐slip faulting. TTZ is imaged as a narrow (10–15 km) transform zone separating the thinned East European cratonic crust from the Małopolska Block.

The updated geological and potential fields data on the East European Platform margin in SE Poland confirm the existence of several regional units differing in Ediacaran to Silurian development: the Upper Silesian Block, Małopolska Block and Łysogóry Block. All the blocks are characterized by a distinct crustal structure seen in Vp velocity models obtained from the seismic refraction data of the CELEBRATION 2000 Programme. The first two units are interpreted as exotic terranes initially derived from Avalonia-type crust and ultimately accreted before the late Early Devonian. The Łysogóry Block is probably a proximal terrane displaced dextrally along the Baltica margin. The sutures between the terranes do not precisely match lateral gradients in Vp models. This is partly explained by a limited resolution of refraction seismic data (20 km wide interpretative window). Most of the difference is related, however, to a post-accretionary tectonism, mainly Variscan transtension–transpression. The latter processes took advantage of lithospheric memory recorded earlier as zones of rheological weakness along the former suture zones. The course of the East European Platform margin (= Teisseyre–Tornquist Zone) corresponds most likely to the Nowe Miasto–Zawichost Fault marking the NE boundary of the proximal Łysogóry Terrane.

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