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A submersible study of the products of a large submarine eruption demonstrates the influence of the ocean on eruption dynamics.

25 26A long-standing conceptual model for deep submarine eruptions is that high hydrostatic pressure 27 hinders degassing and acceleration, and suppresses magma fragmentation. The 2012 submarine 28 rhyolite eruption of Havre volcano in the Kermadec arc provided constraints on critical 29 parameters to quantitatively test these concepts. This eruption produced a > 1 km 3 raft of floating 30 pumice and a 0.1 km 3 field of giant (>1 m) pumice clasts distributed down-current from the vent. 31We address the mechanism of creating these clasts using a model for magma ascent in a conduit. 32We use water ingestion experiments to address why some clasts float and others sink. We show 33 that at the eruption depth of 900 m, the melt retained enough dissolved water, and hence had a 34 low enough viscosity, that strain-rates were too low to cause brittle fragmentation in the conduit, 35 despite mass discharge rates similar to Plinian eruptions on land. There was still, however, 36 enough exsolved vapor at the vent depth to make the magma buoyant relative to seawater. 37Buoyant magma was thus extruded into the ocean where it rose, quenched, and fragmented to 38 produce clasts up to several meters in diameter. We show that these large clasts would have 39 floated to the sea surface within minutes, where air could enter pore space, and the fate of clasts 40 is then controlled by the ability to trap gas within their pore space. We show that clasts from the 41 raft retain enough gas to remain afloat whereas fragments from giant pumice collected from the 42 seafloor ingest more water and sink. The pumice raft and the giant pumice seafloor deposit were 43 thus produced during a clast-generating effusive submarine eruption, where fragmentation 44 occurred above the vent, and the subsequent fate of clasts was controlled by their ability to ingest 45 water. 46 3 47

Sex chromosomes in flowering plants, in contrast to those in animals, evolved relatively recently and only a few are heteromorphic. The homomorphic sex chromosomes of papaya show features of incipient sex chromosome evolution. We investigated the features of paired X- and Y-specific bacterial artificial chromosomes (BACs), and estimated the time of divergence in four pairs of sex-linked genes. We report the results of a comparative analysis of long contiguous genomic DNA sequences between the X and hermaphrodite Y (Y(h)) chromosomes. Numerous chromosomal rearrangements were detected in the male-specific region of the Y chromosome (MSY), including inversions, deletions, insertions, duplications and translocations, showing the dynamic evolutionary process on the MSY after recombination ceased. DNA sequence expansion was documented in the two regions of the MSY, demonstrating that the cytologically homomorphic sex chromosomes are heteromorphic at the molecular level. Analysis of sequence divergence between four X and Y(h) gene pairs resulted in a estimated age of divergence of between 0.5 and 2.2 million years, supporting a recent origin of the papaya sex chromosomes. Our findings indicate that sex chromosomes did not evolve at the family level in Caricaceae, and reinforce the theory that sex chromosomes evolve at the species level in some lineages.

Improved constraints on the mechanical behavior of magma chambers is essential for understanding volcanic processes; however, the role of crystal mush on the mechanical evolution of magma chambers has not yet been systematically studied. Existing magma chamber models typically consider magma chambers to be isolated melt bodies surrounded by elastic crust. In this study, we develop a physical model to account for the presence and properties of crystal mush in magma chambers and investigate its impact on the mechanical processes during and after injection of new magma. Our model assumes the magma chamber to be a spherical body consisting of a liquid core of fluid magma within a shell of crystal mush that behaves primarily as a poroelastic material. We investigate the characteristics of time‐dependent evolution in the magma chamber, both during and after the injection, and find that quantities such as overpressure and tensile stress continue to evolve after the injection has stopped, a feature that is absent in elastic (mushless) models. The time scales relevant to the postinjection evolution vary from hours to thousands of years, depending on the micromechanical properties of the mush, the viscosity of magma, and chamber size. We compare our poroelastic results to the behavior of a magma chamber with an effectively viscoelastic shell and find that only the poroelastic model displays a time scale dependence on the size of the chamber for any fixed mush volume fraction. This study demonstrates that crystal mush can significantly influence the mechanical behaviors of crustal magmatic reservoirs.