Ancient genomes reveal structural shifts after the arrival of Steppe-related ancestry in the Italian Peninsula Highlights d 22 genomes from Northeastern and Central Italy dated between 3200 and 1500 BCE d Arrival of Steppe-related ancestry in the central Italian Peninsula by 1600 BCE d Close patrilineal kinship patterns within commingled Chalcolithic cave burials d Roman Imperial period had a stronger effect on phenotype shifts than the Bronze Age

The mobility patterns in the Italian peninsula during prehistory are still relatively unknown. The excavation of the Copper Age and Bronze Age deposits in La Sassa cave (Sonnino, Italy) allowed to broaden the knowledge about some local and regional dynamics. We employed a multi-disciplinary approach, including stable (carbon and nitrogen, C and N, respectively) and radiogenic (strontium, Sr) isotopes analyses and the identification of the cultural traits in the material culture to identify mobility patterns that took place in the region. The Sr isotopic analyses on the human bones show that in the Copper Age and at the beginning of the Bronze Age, the cave was used as a burial place by different villages, perhaps spread in a radius of no more than 5 km. Stable isotopes analyses suggest the introduction of C4 plants in the diet of the Middle Bronze Age (MBA) communities in the area. Remarkably, in the same period, the material culture shows increasing influxes coming from the North. This evidence is consistent with the recent genomic findings tracing the arrival of people carrying a Steppe-related ancestry in Central Italy in MBA.

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<p><strong>Abstract.</strong> The survey of ancient cave can generally be performed by traditional topographic methods that allow also its georeferencing in a global reference frame; some difficulties may arise when there are narrow tunnels that do not consent the use of a total station or a terrestrial laser scanner. In such cases a visual-based approach can be used to produce, both the followed path and the 3D model of the hypogeal environment. A prompt photogrammetric survey has been used to reconstruct the morphology of the La Sassa Cave, situated in the municipality of Sonnino (Latina), in the lower Lazio region. In this cave, a very large quantity of Pleistocene animal bones was found, together with several fragments of Copper Age human bones and Bronze Age impasto potsherds.</p><p> The survey was carried out using a DSLR full frame camera Nikon D800E with a Nikkor 16<span class="thinspace"></span>mm fisheye lens pre-calibrated. During the acquisition, several targets were measured in order to contain the deformations model. The photogrammetric model has been georeferenced using 3 GCPs positioned outside the cave entrance where a double frequency GNSS receiver has acquired data in static session mode.</p>