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Paleoenvironmental variations that occurred in Italy from the Middle Pliocene to the Late Pleistocene are described. Th e number of large mammal species seems increased moderately, especially from the Galerian to the Late Aurelian biochronological units. On the contrary, the paleobotanical data show a decrease of the forest cover from the Middle Pliocene to the late Early Pleistocene and an increase of lands occupied by prairies and steppes. Th is change is also supported by the appearance of hypsodont taxa among mammals. Th e distribution of mammal faunas between the Adriatic and Tyrrhenian sides of Central Italy, during the Middle and Late Pleistocene, seems not to be infl uenced by climatic and environment diff erences. When the Adriatic data are more complete, it is possible, in fact, to observe a trend that is fairly close to that of the Tyrrhenian. Th e majority of megaherbivorous taxa has an Asian origin, and it can be hypothesized that in the interglacial phases, the Tosco-Emilian Apennines allowed the taxa coming from the northeast to enter and spread out into the more diversifi ed Tyrrhenian side, whereas during the glacial periods the

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The Aurelian Mammal Age for peninsular Italy was introduced on the basis of faunal assemblages mainly recovered at sites along the Via Aurelia west of Rome. These sites exposed a set of sedimentary deposits currently attributed to the Aurelia and to the Vitinia Formations correlated with MIS 9 and MIS 7, respectively. In the present paper we reconstruct the geologic-stratigraphic setting in the western sector of Rome within the wider context of glacio-eustatically controlled, geochronologically constrained aggradational successions defined for this region. We present a chronostratigraphic study based on dedicated field surveys, that, combined with five new40Ar/39Ar ages and eighteen trace-element and EMP glass analyses of volcanic products, allow us to revise age and correlation with the Marine Isotopic Stages for 10 sites out of 12 previously attributed to the Aurelia Formation and the Torre in Pietra Faunal Unit. In particular, we demonstrate a MIS 13/MIS 11 age for several sections along the Via Aurelia between Malagrotta and Castel di Guido. Based on this new geochronological framework, the first occurrences of Canis lupus and Vulpes vulpes in Italy are antedated to MIS 11, within the Fontana Ranuccio Faunal Unit of the Galerian Mammal Age, consistent with the wider European context. This contribution is intended as the groundwork for a revision of the Middle Pleistocene Mammal Ages of the Italian peninsula, according to the improved chronostratigraphy of the geologic sections hosting the faunal assemblages