A scheme of phylogeny in the Proteaceae is derived from analysis of new and previously available data on morphological, anatomical and chromosomal characters. The probable attributes of a common ancestor are indicated and it is concluded that the family has no close relatives, although it possibly diverged early from the Rosiflorean line. Chromosomal evolution has been complex, with early polyploidy, repeated dysploid reductions and great change in chromosome size, from a probable base of x = 7. Habit, wood anatomy, leaf form and sequence, chemical features, inflorescences, pollination and pericarp anatomy are discussed in relation to adaptation, phylogeny, parallelism and convergence. Distribution is analysed ecogeographically and in relation to the fossil record and chronology of continental movement. Ancestors of major groups may have differentiated before the break‐up of Gondwanaland. Early members of almost all tribes were probably trees of mesothermic closed forests; xeromorphic characters found in many modern genera are secondary and of multiple origin in Australia, South Africa, New Caledonia and to a lesser degree in South America. The Proteaceae, with 75 genera, are divided into 5 subfamilies (3 new) comprising 14 tribes, further subdivided into 33 units of subtribal level. New genera described are Toronia, Acidonia, Pycnonia, Triunia, Malagasia, Athertonia, Virotia, Floydia.

185Hill, KD. & Johnson, L.A.S. (National Herbarium of New South Wales, Royal Botanic Gardens, Sydney, Australia 2000) 1995. Systematic studies in the eucalypts. 7. A revision of the bloodwoods, genus Corymbia (Myrtaceae). Telopea 6(2-3): 185-504. A taxonomic revision of the bloodwoods (including 'ghost gums') is presented. Previous cladistic studies in the eucalypts are re-evaluated; new phylogenetic analyses are presented for the Angophora + Eucalyptus sens. lat. group, the Angophora + blood wood dade, and the sections and series of the 'red bloodwoods'.The non-angophoroid, non-bloodwood eucalypts are discussed, but a formal taxonomic system for these is not presented here. The bloodwoods are placed in a new genus Corymbia K.D. Hill & L.A.S. Johnson. Relationships within the group are examined, and morphological interpretation and phytogeography are discussed. An infra generic classification is presented, with keys to infra generic groupings and species. One hundred and thirteen species are described, 33 of them new (c. chlorolampra, C. maritima, C. ligans, C. plena, C. hendersonii, C. rubens, C. porphyritica, C. semiclara, C. tum esc ens, C. opacula, C. pedimontana, C. sphaerica, C. chartacea, C. dunlopiana, C. pachycarpa, C. papillosa, C. catenaria, C. paracolpica, C. arafurica, C. bella, C. disjuncta, C. pauciseta, C. karelgica, C. dendromerinx, C. torta, C. chillagoensis, C. inobvia, C. blakei, C. candida, C. punkapitiensis, C. aparrerinja, C. flavescens, C. paractia). New subspecies are described or recognised in C. trachyphloia (subspp. amphistomatica and carnarvonica), C. arnhemensis (subsp. monticola), C. hylandii (subsp. peninsularis), C. eremaea (subsp. oligocarpa), C. cadophora (subsp. pliantha), C. setosa (subsp. pedicellaris), C. ferruginea (subsp. stypophylla), C. pachycarpa (subsp. glabrescens), C. papillosa (subsp. globifera), C. deserticola (subsp. mesogeotica), C. torta (subspp. allanii and mixtifolia), C. blakei (subsp. rasilis), C. candida (subspp. dipsodes and lautifolia), C. ferriticola (subsp. sitiens), C. grandifolia (subspp. lamprocardia and longa). A new name is provided for the species illegitimately known asEucalyptus perfoliata (c. cadophora), and a valid publication of C. aparrerinja (published invalidly as a variety of E. papuana by Blakely) is effected. Two species are reinstated from synonymy (c. polysciada and C. variegata), and two raised from infra specific to specific rank (c. dallachiana and C. dimorpha). Lectotypes are designated for the names E.

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A new classification is presented of all taxa of EuaaZyptus (and Angophora) (Myrtaceae Leptospermoideae), on the basis of studies from many disciplines and extensive field experience. This is not in the traditional revisionary form and formal nomenclatural innovations at the species and subspecies level will follow later. Infrageneric classification into subgenera, sections, series, and subseries follows a rationalised plan explicitly divorced from the traditional system embodied in the International Code of Botanical Nomenclature. This is accompanied by an equivalent and flexible system using 1-to 6-letter coded designations for taxa of the various ranks, which embodies the whole classificatory structure. There is a comprehensive index to specific and infraspecific names. Discussion covers the kinds of evidence used, the inflorescence, the operculum, the ovule, and the seed, as well as genetic behaviour, the range of variation-patterns found, and the case for recognition of segregate genera. Recognition of two (only) such genera (EuaaZyptus s. str. and Symphyomyrtus) as proposed by some recent authors is considered oversimplified and contrary to the evidence.Although division into a number of genera may perhaps be desirable in the future, it seems best at present to consider all the eucalypts as constituting a single genus with eight subgenera.Angophora would logically be included as one of these but, to avoid possible future reversals, its generic status •is not • formally reduced at this• stage; IX L 1.

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