Lassa fever virus (LAS), the most dangerous human pathogen among the Arenaviridae, has two single-stranded RNA genomic segments termed large (L) and small (S) (Lukashevich et al., 1984). The LAS S RNA (3n4 kb) encodes the nucleoprotein (NP) and the envelope glycoprotein (GP) in an ambisense coding arrangement (Clegg & Oram, 1985 ;
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