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Background: Type II sucrose transporters (SUTs) are more selective than type I SUTs. Results: OsSUT1 was modified using gene shuffling. Variants were selected for the ability to transport the fluorescent ␤-glucoside esculin. Conclusion: Replacement of five amino acids conferred substrate specificity typical of type I SUTs. Significance: The first loop and top of TMS2 and TMS5 are involved in controlling substrate specificity in SUTs.

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