Hindeodus parvus and Hindeodus typicalis occur in a deep-water chert and claystone section in the Mino Terrane, Japan, which has been identified as a Jurassic accretionary complex. Conodont fossils are preserved as natural assemblages of impression fossils on bedding planes in claystone. In this study, 13 assemblages of Hindeodus species were recognized, comprising at most 13 elements which generally maintain the original composition and structure of an apparatus. We discriminated pairs of carminiscaphate P 1 , angulate P 2 and makellate M elements, as well as a single alate S 0 element and two digyrate and four bipennate elements constituting the S array. Although the digyrate and bipennate elements are preserved in the S 2 and S 3-4 positions, respectively, a pair of S 1 elements was not found due to incompleteness in the natural assemblages. The conodont biostratigraphy indicates that the lithological boundary between chert and claystone units in the study section corresponds exactly to the Permian-Triassic boundary.
Moderately well-preserved Late Permian to Middle Triassic radiolarians are identified in chert beds that occur in the Shan-Thai Block of northern Thailand. These radiolarians are identical to the faunas of the Late Permian Neoalbaillella ornithoformis and N. optima Assemblage Zones and the Triassic Parentactinia nakatsugawaensis and Triassocampe coronata Assemblage Zones reported in chert sequences of Japan. We discovered the radiolarian faunas, apparently indicating Late Permian and Early Triassic ages, in almost continuous sequences of chert and shale exposed in the north of Chiang Mai. The occurrence of these radiolarian faunas provides important data to solve the P/T (Permian/Triassic) boundary in pelagic sequences. Our present discovery also furnishes significant data to reconstruct the paleobiogeography of Mainland Thailand during Late Permian to Middle Triassic times. Fifty species belonging to 35 genera, including three unidentified genera, are investigated taxonomically. Four new species Pseudospongoprunum? chiangdaoensis, Cenosphaera igoi, Cenosphaera? rugosa, and Tlecerina? apsornae are described.
The apparatus of an Early Triassic conodont Neostrachanognathus tahoensis Koike, 1998 from Oritate, Kumamoto Prefecture, Japan, and a species of Neostrachanognathus from Oman were reconstructed. On the basis of five natural assemblages from the Oritate area, the threedimensional apparatus model of N. tahoensis is interpreted as bilaterally symmetrical and composed of 14 elements consisting of pairs of P 1 , P 2 , P 3 , S 1 , S 2 , S 3 , and S 4 elements. The P 1 and P 2 elements are coniform elements, the P 3 elements are digyrate forms, and the S elements are bipennate ramiforms. The S elements are arranged rostrally in the apparatus and the pairs of the P 1 , P 2 , and P 3 elements are subvertically arranged caudally and ventrally to the S array. One of the natural assemblages was formed by rostrocaudal collapse of the apparatus on the sea floor, whereas the other assemblages indicate that conodont animals came to rest nearly parallel with the substrate prior to burial. A collection of isolated elements from Jabal Safra, Oman, includes a second species of Neostrachanognathus with a comparable apparatus.
The Middle and Upper Ordovician sequence of the Langkawi Islands, northwestern peninsular Malaysia, contains 20 species of conodonts belonging to 15 genera and four unidentified species, which are described and illustrated. The following four biostratigraphic zones are established for the study area: the Scolopodus striatus assemblage zone, the Periodon sp. A range zone, the Baltoniodus alobatus range zone, and the Hamarodus europaeus range zone, in ascending order. The Middle Ordovician fauna belongs to the low-latitude, warm-water Australian Province. Conodonts of the H. europaeus zone represent the HDS (Hamarodus europaeus-Dapsilodus mutatus-Scabbardella altipes) biofacies, which has been reported from the cool-water North Atlantic Faunal Region. The middle Arenigian limestones in the study area were deposited on a shallow-water shelf, whereas the late Arenigian to middle Darriwilian limestones formed in hemipelagic deeper-water conditions on an outer shelf or slope.
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