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Light and dark bands in fully recrystallized fossil hermatypic corals are generally interpreted to represent annual growth increments reflecting a photosymbiotic life style-an interpretation of far reaching significance in palaeoecology. In this paper we describe annual growth bands in the colonial coral Porites in a perfect (aragonite and microstructures retained) and fully recrystallized (sparry calcite mosaic) style of preservation from sediments of Late Miocene age (Crete, Greece). Analysis of a continuous spectrum of transitional preservational stages shows that in Miocene Porites preservation of the growth banding was controlled by preferential dissolution of the highdensity band associated with cementation by drusy calcite spar during freshwater diagenesis/shallow burial diagenesis. Marine precipitates (pelletoidal Mg-calcite) preferentially accumulated along tabulate dissepiments producing an additional growth rhythmicity. Massive Porites had annual growth rates of ∼4.0 mm, whereas in ramose branching Porites, a conspicuous banding is formed by concentrations of marine micropelletoidal cement along dissepiments at ∼1.8 mm spacing. If taken as annual growth increments, these bands represent very low extension rates, however, they may rather reflect subannual forcing functions (i.e., lunar cycles). An identical scenario of precipitation and concentration of pelletoidal carbonate along dissepiments and dissolution-controlled documentation of growth bands can be inferred for Late Jurassic microsolenids. Therefore, growth bandings in fossil corals potentially reflect both, monthly and annual cycles. Consequently, care must be taken when using coral growth bands in palaeoecology and palaeoclimatology.

In modern oceans, the transition zone between the tropical and temperate carbonate province is gradual and covers a wide latitudinal belt. Little knowledge exists regarding the geological signatures of this zone. This paper describes a late Miocene (early Tortonian-early Messinian) transitional carbonate system that combines elements of the tropical and cool-water carbonate systems (Iraklion Basin, island of Crete, Greece). As documented in stratal geometries, the submarine topography of the basin was controlled by tilting blocks. Coral reefs formed by Porites and Tarbellastrea occurred in a narrow clastic coastal belt along a central Cretan landmass and steep escarpments formed by faulting. On the gentle dip-slope ramps of those blocks having the widest geographical distribution within the basin, extensive covers of level-bottom communities existed in a low-energy environment. Isolated colonial corals were present in the shallow segments of the ramps. Consistent patterns of landward and basinward shift of coastal onlap in all outcrop studies reveal an overriding control of third-and fourthorder sea-level changes on sediment dynamics and facies distributions over block movements. An increasingly dry climate and the complex submarine topography of the fault-block mosaic kept sediment and nutrient discharge from a central Cretan landmass at a minimum. The skeletal limestone facies therefore reflects oligotrophic conditions and sea surface temperatures near the lower threshold temperature of coral reefs in a climatic position transitional between the tropical coral reef belt and the temperate zone. It is suggested that the recognition of an overall late Miocene aridification trend helps to explain the Mediterranean-wide distribution of shallow-marine carbonates, both cool-water and warm-water, in settings adjacent to uplifting mountain ranges (intramontane basins).

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