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Wood-pastures are threatened biotopes in which trees and livestock grazing maintain high conservation values. However, browsing may threaten tree regeneration, whereas abandonment leads to tree encroachment. We studied the regeneration of trees in a grazed and abandoned boreal wood-pastures. In grazed sites, the density of young spruces (Picea abies) was high, while the density of young birches (Betula spp.) was very low. Sprucification can be prevented only by removing spruces. The number of young birches and pines (Pinus sylvestris) was correlated with the number of junipers (Juniperus communis), probably because thorny junipers protect palatable seedlings from browsing. In abandoned sites, deciduous trees and spruces regenerated abundantly. In the long term, both grazing and abandonment lead to changes in tree species compositions and low diversity wood-pastures. Landscape scale planning and disturbance dynamics are needed for the creation of new wood-pastures and the maintenance of all pasture types within the landscape.

Traditional rural biotopes such as wood-pastures are species-rich environments that have been created by low-intensity agriculture. Their amount has decreased dramatically during the 20th century in whole Europe due to the intensification of agriculture. Wood-pastures host some fungal species that prefer warm areas and are adapted to semi-open conditions, but still very little is known about fungi in these habitats. We studied how management, historical land-use intensity, present grazing intensity, time since abandonment, and stand conditions affect the species richness and community composition of ectomycorrhizal fungi. We surveyed fruit bodies on three 10 m x 10 m study plots in 36 sites and repeated the surveys three times. Half of the sites were currently unmanaged but had a grazing history. We measured soil pH,

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