L eaf-cutter ant nests have a complex and specialized internal structure that allows them not only to maintain stable microenvironmental conditions where they can shelter their colonies, but also to protect themselves from various predators and intruders (Bollazzi and Roces 2002;Diehl-Fleig and Diehl 2007;Velásquez-Múnera et al. 2008). These characteristics allow colonies to have multiple types of interactions with other species, varying from those that can use colony debris to others that present coevolutionary relationships (Mehdiabadi
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