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During the Palaeocene and Eocene Epochs (65.5-33.9 Ma) the Earth experienced the warmest conditions of the Cenozoic. This Palaeogene greenhouse episode is characterized by several short-lived negative carbon isotope (ä 13 C) excursions, which are usually interpreted as transient warming events ('hyperthermals') as indicated by rising temperatures of surface and bottom waters, and accompanied carbonate dissolution in deep-sea settings.

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Danian (lower Paleocene) sequences in north-central Tunisia are dominantly composed of marls and shales but a conspicuous, indurated glauconite-bearing marker bed is associated with the P3a-P3b transition. This glauconite bed is considered to correlate with the Latest Danian Event (LDE) described from the Nile Basin in Egypt, with the 'top Chron C27n event' (Atlantic and Pacific Oceans) and with the 'CIE-DS1' (Zumaia, Spain). The LDE is thought to reflect a short period of global warming, similar to the Paleocene Eocene Thermal Maximum, but of lesser magnitude. The presence of a glauconitic bed at the P3a/P3b transition in Tunisia suggests that the sequence is condensed, which is confirmed by planktic foraminifer and nannofossil biostratigraphy, and by the absence of the typical LDE beds found in most Egyptian sections. Benthic foraminiferal assemblages were quantitatively analyzed in three sections in north-central Tunisia (Sidi Nasseur, Garn Halfaya, Elles) in order to characterize the paleoenvironmental evolution during the late Danian and compare this with sections in Egypt. The benthic foraminiferal assemblages indicate that the Tunisian sections were located along a depth-and a paleoenvironmental gradient on the shelf north of Kasserine Island. The reconstructed depth range is comparable with sections in Egypt, and encompasses middle neritic (Sidi Nasseur section) to shallow upper bathyal depths (Elles section, with up to 16% Gavelinella beccariiformis below the P3a/P3b). Like in Egypt, assemblage shifts across the P3a/P3b subzonal transition indicate shallowing and a transition to a more eutrophic paleoenvironment, characterized by relatively high abundances of buliminids and Stainforthia sp. The δ 13 C and δ 18 O records generated on well-preserved specimens of the ostracode species Bairdia failed to demonstrate the presence of the carbon isotope excursion (CIE) associated with the LDE at the base of Subzone P3b. A condensed section or a hiatus associated with the glauconite bed would explain why the CIE is not recorded in the Tunisian sections.

Two detailed records (NSF and 05NSC, Sidi Nasseur, Tunisia) across the Danian/Selandian (D/S) boundary were investigated for their micropaleontological content. Calcareous nannofossils and planktic foraminifera provided a biostratigraphic framework. The interval spans part of planktic foraminiferal Zone P2, Subzone P3a and part of Subzone P3b. This corresponds to calcareous nannoplankton Zone NP4. Using a more detailed nannofossil zonation the studied section spans part of Zone NTp6, Zone NTp7a and part of NTp7b. Quantitative ostracod and qualitative benthic foraminiferal data were used to characterize environmental changes across the D/S boundary. The two subsections have yielded a total of 50 ostracod taxa. The ostracod assemblage of the entire section belongs to the Southern Tethyan Type showing subtle but distinct changes up section. Based on statistical analysis of the quantitative ostracod data, faunal changes at a glauconitic maker bed (P3a/P3b boundary) were demonstrated. The local Reticulina proteros assemblage, with the typical species R. proteros, Oertliella vesiculosa and Cytheroptheron lekefense, is gradually replaced by the Protobuntonia nakkadii assemblage, with the typical species Cristaeleberis arabii, Xestoleberis tunisiensis, Cytheropteron sp. and P. nakkadii, across the glauconitic bed. The benthic foraminifera also demonstrated distinct changes at this marker bed. The changes in ostracods and foraminifera are related to changes in paleoproductivity and an overall relative sea-level fall. The lithological and faunal changes at the P3a/P3b zone boundary within the Sidi Nasseur sections seem to correspond to the D/S boundary in the type region in Danmark and are characterized by a significant hiatus, yielding this section not suitable as a GSSP candidate for this boundary.