The subfamily Mutisioideae (74 genera, ca. 865 species), which comprises three tribes, Mutisieae (43 genera, ca. 500 species), Nassauvieae (25 genera, ca. 315 species), and Stifftieae (six genera, 48 species), is analyzed at the generic level. A total of 87 genera traditionally related to Mutisioideae were studied. The genera of Mutisioideae are described and illustrated, and keys to tribes and genera are given. Corollas, anthers, styles, and pollen, are analyzed and discussed. The styles (smooth, rugulose to papillose, papillae short and rounded) constitute the main characters for delimiting the subfamily. The presence and distribution of the style papillae support the recognition of three tribes, although other characters as corolla shape contribute to their definition. The pollen of Nassauvieae can be clearly differentiated from the pollen of Mutisieae and Stifftieae. The morphology supports the exclusion of Brachylaena, and Warionia but not the exclusion of other taxa, for example, Hecastocleis, the Gochnatia complex, or the Ainsliaea group members.Resumen La subfamilia Mutisioideae (74 géneros, ca. 865 especies), que comprende tres tribus, Mutisieae (43 géneros, ca. 500 especies), Nassauvieae (25 géneros, ca. 315 especies), y Stifftieae (6 géneros, 48 especies), es analizada a nivel genérico. Se estudió un total de 87 géneros que tradicionalmente fueron relacionados a Mutisioideae. Se describen e ilustran los géneros de Mutisioideae, y se proveen claves para las tribus y los géneros. Se analizan y discuten las corolas, anteras y estilos. Los caracteres del estilo (glabro, ruguloso a papiloso, las papilas cortas y redondeadas) son los más importantes para delimitar la subfamilia. La presencia y distribución de las papilas del estilo ayudan al reconocimiento de las tres tribus, aunque otros caracteres como la forma de la corola contribuyen a su definición. El polen de Nassauvieae se puede diferenciar claramente del polen de Mutisieae y Stifftieae. La morfología apoya la exclusión de Brachylaena,

Athroismeae is a small tribe of the Asteraceae‐Asteroideae, the members of which show considerable variation in morphology. A molecular phylogenetic study of the tribe is presented for the first time, based on plastid (ndhF, trnH‐pssbA, trnL‐trnF) and nuclear data (ETS, ITS). The phylogenetic relationships between the different genera within Athroismeae are discussed, and in addition, three unispecific genera: Anissochaeta, Artemissiopssiss and Symphyllocarpuss as well as Duhaldea (Inula) sstuhlmannii, all earlier placed in other tribes, are here shown to belong within Athroismeae. Symphyllocarpuss is sister to Centipeda and the earlier Symphyllocarpinae includes Centipedinae in synonymy. Furthermore, Cardossoa and Philyrophyllum are found to be integrated within Anissopappuss and their generic status cannot be maintained. An outline of an amended circumscription of the Athroismeae is presented, with three new combinations and a description of the new subtribe Lowryanthinae.

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A new genus from Venezuela, Tuber¬ culocarpus, is described and illustrated, and the combination T. ruber is made. Thelechitonia and Complaya are placed in synonymy of the formerly unispecific Sphagneticola, and a lectotype is chosen for the generitype S. ulei and for Verbesina calendulacea. Sphagneticola includes cultivated Wedelia trilobata, and the combinations S. brachycarpa, S. calendulacea, S. gracilis, and S. trilobata are made. Wulffia is a taxonomic synonym of Tilesia, and com¬ binations in Tilesia are proposed for the three spe¬ cies and one variety formerly placed in Wulffia: Ti¬ lesia baccata, T. macrocephala, T. rubens, and T. baccata var. discoidea. The basionym of the most widespread species of Tilesia (Coreopsis baccata) has previously been attributed to Linnaeus filius, but was published earlier by Linnaeus. Elaphandra moriana from French Guiana is described, and the combi¬ nation Elaphandra lehmannii is made. Oyedaea wurdackii is described from Venezuela and is illustrated, and the combination Oyedaea tepuiana is proposed. The combination Verbesina ligulata is provided, and V. guianensis is lectotypified and is an earlier name for V. schomburgkii. Elaphandra sucrensis is reduced to synonymy of E. verbesinoides, E. lucidula is re¬ duced to E. ulei, Oyedaea blakeana is reduced to 0. scaberrima, and Wulffia trujilloi is reduced to Tilesia macrocephala. Volume 6, Number 4 1996 Pruski Guayana Highland Compositae 405 when present (one species is discoid), by ovate eglandular anther appendages, these commonly abaxially black though sometimes tan within or distally, by erect or laxly recurved (not strongly coiled) papillose style branches, by rostrate (necked) cypselas that have a reduced or aristate pappus, and by lack of obvious carpopodia and elaiosomes.

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