We demonstrate experimentally that a passerine exploits tail spreading to intercept the downward flow induced by its wings to facilitate the recovery of its posture. The periodic spreading of its tail by the White-eye bird exhibits a phase correlation with both wingstroke motion and body oscillation during hovering flight. During a downstroke, a White-eye's body undergoes a remarkable pitch-down motion, with the tail undergoing an upward swing. This pitch-down motion becomes appropriately suppressed at the end of the downstroke; the bird's body posture then recovers gradually to its original status. Employing digital particle-image velocimetry, we show that the strong downward flow induced by downstroking the wings serves as an external jet flow impinging upon the tail, providing a depressing force on the tail to counteract the pitch-down motion of the bird's body. Spreading of the tail enhances a rapid recovery of the body posture because increased forces are experienced. The maximum force experienced by a spread tail is approximately 2.6 times that of a non-spread tail.

The flight of a small bird under the influence of the ground effect is numerically investigated with a complete three-dimensional model including the bird's body and wings. The flight mode is not the conventional steady gliding flight but an unsteady flight consisting of flapping, twisting, and folding motions. As the bird approaches the ground, the average lift force gradually increases while the average drag force decreases. At a particular distance, the average lift force increases by approximately 47%, whereas the average drag force decreases by nearly 20%, relative to the absence of the ground effect. Because of the ground, the improved aerodynamic performance in flapping flight is much more significant than in steady flight, in which the modification of the lift-drag ratio is typically less than 10%. On the basis of the flow field, regardless of the presence or absence of the ground, there exists no evidence for an obstruction of a wing-tip vortex, which is a remarkable phenomenon and accounts for the improved performance in steady flight. The extent of the region of high pressure beneath the wing in the near-ground case seems to surpass that in the far-ground case, accounting for the greater lift and thrust forces in the near-ground case. This air cushion beneath the wing, known as the cram effect, is the dominant factor of the ground effect on a flapping bird.

Some small birds typically clap their wings ventrally, particularly during hovering. To investigate this phenomenon, we analyzed the kinematic motion and wake flow field of two passerine species that hover with the same flapping frequency. For these two birds, the ventral clap is classified as direct and cupping. Japanese White-eyes undertake a direct clap via their hand wings, whereas Gouldian Finches undertake a cupping clap with one wing overlaying the other. As a result of their morphological limitation, birds of both greater size and wing span cup their wings to increase the wing speed during a ventral clap because of the larger wing loading. This morphological limitation leads also to a structural discrepancy of the wake flow fields between these two passerine species. At the instant of clapping, the direct clap induces a downward air velocity 1.68 times and generates a weight-normalized lift force 1.14 times that for the cupping clap. The direct clap produces a small upward jet and a pair of counter-rotating vortices, both of which abate the transient lift at the instant of clapping, but they are not engendered by the cupping clap. The aerodynamic mechanisms generated with a ventral clap help the small birds to avoid abrupt body swinging at the instant of clapping so as to maintain their visual stability during hovering.

We provide physical insight into how a small hovering bird attains stabilized vision during downstroke. A passerine generates a lift force greater than its body weight during downstroke, leading to a substantial swing of the bird body, but the bird's eyes are nearly stable. Employing digital particle-image velocimetry, we demonstrate that a hovering passerine generates a lift force acting dorsal to the center of mass, concurrently resulting in rotational and translational displacements of the bird's body. The most notable finding is that the rotational and translational displacements at the bird's eyes almost cancel each other; the displacement of the eye is ~8% that of the trailing tip of the tail. This aerodynamic trick enables a bird to attain stabilized vision beneficial for the inspection of the environment.

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