The upper plate deformation pattern reflects the mechanical behavior of subduction zones. Here we focus on the consequences of the entrance of a buoyant bank into the Caribbean subduction zone during the Eocene by studying the oldest exposed rocks belonging to the Lesser Antilles volcanic arc. Using a novel geochronological data set, we show that the volcanic arc activity on the island of St. Barthelemy spanned over the mid‐Eocene to early Miocene with a westward migration of the tectono‐volcanic activity, which is comparable to what has already been observed on other volcanic islands in the Lesser Antilles. The kinematics analysis allows us to identify a switch in the stress field from pure to radial extension at the Oligo‐Miocene hinge with a subhorizontal σ3 that has a mean trend of N20°. A three‐step restoration of the regional deformation indicates that this switch from pure parallel‐to‐the‐trench extension to radial extension may reflect a strain partitioning initiation affecting the upper Caribbean Plate in response to trench bending that followed the entrance of the Bahamas Bank into the subduction zone. We show that the northern end of the Lesser Antilles arc shows a tectono‐volcanic evolution which is similar to the southern one. The north‐south dichotomy in the perpendicular‐to‐the‐trench extension, 15% in the north versus 30% in the south, may reflect different slab ends that are highly curved to the north (restraining the extension in the upper plate) versus a tear to the south (allowing a larger amount of extension within the upper plate).

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Our study aims to reconstruct the palaeogeography of the northern part of the Lesser Antilles in order to analyse whether emerged areas might have existed during the Cenozoic, favouring terrestrial faunal dispersals between South America and the Greater Antilles along the present-day Lesser Antilles arc. The stratigraphy and depositional environments of the islands of Anguilla, St Martin, Tintamarre, St Barthélemy, Barbuda and Antigua are reviewed in association with multichannel reflection seismic data acquired offshore since the 80's in the Saba, Anguilla and Antigua Banks and in the Kalinago Basin, including the most recent academic and industrial surveys. Seven seismic megasequences and seven regional unconformities are defined, and calibrated from deep wells on the Saba Bank and various dredges performed during marine cruises since the 70's in the vicinity of the islands. Onshore and offshore correlations allow us to depict an updated and detailed sedimentary organisation of the northern part of the Lesser Antilles from the late Eocene to the late Pleistocene. Paleogeographic reconstructions reveal sequences of uplift and emergence across hundredswide areas during the late Eocene, the late Oligocene, the early middle-Miocene and the latest Miocene-earliest Pliocene, interspersed by drowning episodes. The ∼200 km-long and ∼20 km-wide Kalinago Basin opened as an intra-arc basin during the late Eocene -early Oligocene. These periods of emergence may have favoured the existence of episodic mega-islands and transient terrestrial connections between the Greater Antilles, the Lesser Antilles and the northern part of the Aves Ridge (Saba Bank). During the Pleistocene, archipelagos and mega-islands formed repeatedly during glacial maximum episodes.

The Grenada Basin is bounded to the east by the active Lesser Antilles Arc, to the west by the north-south trending Aves Ridge, commonly described as a Cretaceous-Paleocene remnant of the "Great Arc of the Caribbean" (Burke, 1988), and to the south by the transpressional plate boundary with South America (Figure 1). This setting led previous authors to propose various models for the origin of the Grenada Basin, most of them assuming the basin to be at least partly floored by oceanic crust that was formed during the

Intriguing latest Eocene land-faunal dispersals between South America and the Greater Antilles (northern Caribbean) has inspired the hypothesis of the GAARlandia (Greater Antilles Aves Ridge) land bridge. This landbridge, however, should have crossed the Caribbean oceanic plate, and the geological evolution of its rise and demise, or its geodynamic forcing, remain unknown. Here we present the results of a land-sea survey from the northeast Caribbean plate, combined with chronostratigraphic data, revealing a regional episode of mid to late Eocene, trench-normal, E-W shortening and crustal thickening by *25%. This shortening led to a regional late Eocene-early Oligocene hiatus in the sedimentary record revealing the location of an emerged land (the Greater Antilles-Northern Lesser Antilles, or GrANoLA, landmass), consistent with the GAARlandia hypothesis. Subsequent submergence is explained by combined trench-parallel extension and thermal relaxation following a shift of arc magmatism, expressed by a regional early Miocene transgression. We tentatively link the NE Caribbean intra-plate shortening to a well-known absolute and relative North American and Caribbean plate motion change, which may provide focus for the search of the remaining connection between 'GrANoLA' land and South America, through the Aves Ridge or Lesser Antilles island arc. Our study highlights the how regional geodynamic evolution may have driven paleogeographic change that is still reflected in current biology.

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Worldwide, forearc trench-perpendicular basins are interpreted as the result of trench-parallel extension possibly due to either strain partitioning as at the Aleutians (Ryan & Scholl, 1989) and Ryukyu (Nakamura, 2004) Subduction Zones, and/or to increasing margin curvature as at the Marianas (Heeszel et al., 2008) and Hellenic trenches (Angelier, 1978; Mascle & Martin, 1990). In more extreme cases, widespread deformation of forearc domains results from the collision of buoyant crustal features (e.g., oceanic plateaus, seamount chains, or continental fragments) which is prone to generate bending and rotation of subduction zones (e.g., Vogt et al., 1976). Strongly curved convergent plate boundaries are subject to alongstrike variations in subduction obliquity and thus commonly associated with large-scale rigid body rotation