Pseudoscorpions form phoretic associations with a wide range of arthropods, including at least 44 families of insects and three families of arachnids.The present work brings up to date phoretic associations between pseudoscorpions and different arthropods and discusses the long-standing controversy over whether this behavior is basically a predatory response or adapted solely for dispersal. That phoresy in pseudoscorpions is of long standing and obligatory in many cases is demonstrated by its continuance for millions of years, as shown by the fossil record. A member of the Chthoniidae attached to a moth in Baltic amber is reported for the first time.
Three species [Ceratinella playa new species, Erigone denticulata Chamberlin & Ivie, Mermessus denticulatus (Banks)] in the spider family Linyphiidae were associated with emergent vegetation in playa wetlands in the Southern High Plains of Texas. Playa wetlands are being rapidly degraded in the Great Plains of the U.S.A. We describe and illustrate the new species Ceratinella playa from a male and female from Briscoe County, Texas. Erigone denticulata is redescribed and illustrated from males and females from Texas. New distributional records are provided for E. denticulata from the U.S.A.: Colorado, Idaho, Nevada, Oregon, Texas, Utah, and Wyoming. All of these except those from Wyoming are new state records. A new county record of Mermessus denticulatus is reported from Texas. RESUMENTres especies [Ceratinella playa nueva especie, Erigone denticulata Chamberlin & Ivie y Mermessus denticulatus (Banks)] de la familia Linyphiidae fueron asociadas a la vegetación emergente en cuerpos de aguas temporales "playas" de humedales en las Grandes Planicies del sur de Tejas. Los humedales están siendo degradados rápidamente en las Planicies de los E.E.U.U. Se describe e ilustra macho y hembra de la nueva especie Ceratinella playa del condado Briscoe, Tejas. Machos y hembras de Erigone denticulata de Tejas son redescritos e ilustrados. Nuevos datos distribucionales son reportados para E. denticulata para los estados de Colorado, Idaho, Nevada, Oregón, Tejas, Utah y Wyoming. Todos, excepto los de Wyoming, son nuevas citas para estos Estados. Mermessus denticulatus se cita por primera vez para Tejas.
A phylogenetic analysis of the spider genus Neoleptoneta Brignoli, 1972 is presented based on molecular sequence variation from three genes (mitochondrial cytochrome c oxidase subunit I, nuclear histone H3 and nuclear 28S rDNA) and including exemplars for all North American leptonetid genera except the ecribellate archoleptonetine Darkoneta. Analysis of concatenated data and independent genes using Bayesian, maximum likelihood and parsimony methods failed to recover Neoleptoneta as monophyletic. The genera Archoleptoneta, Appaleptoneta and Calileptoneta are monophyletic and a sister group relationship is supported between Appaleptoneta and Calileptoneta. Morphological data based on a survey of leptonetid genera using scanning electron and compound light microscopy are discussed and traced on the molecular phylogeny. Images for each North American leptonetine genus are provided, including comparison with Asian and European outgroups. Images of the incertae sedis species Leptoneta brunnea Gertsch, 1974 and Leptoneta sandra Gertsch, 1974 are provided and their generic placement is re-evaluated. Ancestral state reconstruction is used to assess patterns of cave evolution and shows that most species are descended from troglophilic ancestors and that troglobites have evolved at least nine times independently within the North American Leptonetidae. Neoleptoneta is relimited to include seven species restricted to central Mexico including N. bonita (Gertsch, 1974), N. capilla (Gertsch, 1971), N. delicata (Gertsch, 1971), N. limpida (Gertsch, 1974), N. rainesi (Gertsch, 1971) and N. reclusa (Gertsch, 1971) and to include Leptoneta brunnea, giving the new combination N. brunnea (Gertsch, 1974). The remaining species described in Neoleptoneta are placed in three new genera: (1) Chisoneta, gen. nov. from south-western Texas and Nuevo Leon, Mexico, including the four species C. chisosea (Gertsch, 1974), C. isolata (Gertsch, 1971), C. modica (Gertsch, 1974) and C. pecki (Gertsch, 1971), new combinations; (2) Ozarkia, gen. nov. from Arizona and New Mexico north-east to Arkansas, Alabama and Georgia, including the nine species O. alabama (Gertsch, 1974), O. apachea (Gertsch, 1974), O. archeri (Gertsch, 1974), O. arkansa (Gertsch, 1974), O. blanda (Gertsch, 1974), O. georgia (Gertsch, 1974), O. ivei (Gertsch, 1974), O. novaegalleciae (Brignoli, 1979) and O. serena (Gertsch, 1974), new combinations; and (3) Tayshaneta, gen. nov. from Texas south to Coahuila, Mexico, with the eleven species T. anopica (Gertsch, 1974), T. bullis (Cokendolpher, 2004), T. coeca (Chamberlin & Ivie, 1942), T. concinna (Gertsch, 1974), T. devia (Gertsch, 1974), T. furtiva (Gertsch, 1974), T. microps (Gertsch, 1974), T. myopica (Gertsch, 1974), T. paraconcinna (Cokendolpher & Reddell, 2001), T. uvaldea (Gertsch, 1974) and T. valverdae (Gertsch, 1974), new combinations. Leptoneta sandra Gertsch, 1974 cannot be placed in any North American, European or Asian genus and is thus transferred to the new genus Montanineta, gen. nov., giving the new combination Montanineta sandra (Gertsch, 1974).
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