1999. Morphologie et ultrastructure du pollen des Siparunaceae (Laurales). ± Grana 38: 210±217. ISSN 0017-3134.The pollen morphology of Siparuna, Bracteanthus and Glossocalyx, and the ultrastructure of S. decipiens and G. longicuspis are described. All the species have small, inaperturate (omniaperturate), apolar, radiosymmetric, spherical and echinulate pollen grains. The ectexine consists of three layers: 1) a thin, wavy tectum linking the bases of large spines, with some small spines and rare ®laments; 2) an infratectum with large, subglobose granules continuous with the large spines, connecting the tectum and the foot-layer , called « columellar granules », and mixed with more or less numerous small granules, especially near the tectum; 3) a very thin, but well-de®ned foot-layer with oblique and short lamellae extending into the underlying layer. This inner layer is continuous and granulose and is interpreted as endexine or « endexine-like ». This type of exine represents an intermediate state between columellar and granular struture. The intine is much thicker than the exine and uniform around the pollen grain and consists of three zones in the mature pollen of Glossocalyx in which the inner layer is very thick and channelled. This palynological study con®rms the status of the Siparunaceae as a separate family, but neither supports no contradicts the relationships with the Gomortegaceae.
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