Thymus jenisseensis is endemic to Southern Siberia. The structure of Thymus jenisseensis were studied with the use of an architectural approach. The architectural unit consist of branched compound skeletal axis of the 1st and 2nd order, and is repeated many times in the structure of adults. As a result of studying the architectural units of individuals growing in different sites of a coenopopulation, in the upper border of the forest belt (Tsagan-Shibetu, Republic of Tuva), differences were identified. An architectural unit consisting of branched orthotropic or ascending basisympodially accreting compound skeletal axes develops on a site of a dry riverbed; an architectural unit consisting of branched orthotropic or ascending acrosympodially accreting compound skeletal axes develops on a site of high-altitude steppe on a plain. The diversity of compound skeletal axes in the structure of architectural units contributes to the formation of two biomorphs (dwarf subshrub and dwarf shrub), changes in the vitality and duration of development of T. jenisseensis individuals. The identified features of architecture are morphological mechanisms of adaptation of the species to living conditions.
Seasonal rhythms of plant development are related to plant adaptation to surrounding ecological coenotic and climatic conditions. To discover the patterns of seasonal development of plants in different phytocoenoses, it is essential to observe individual shoots and the entire course of shoot formation. So far the data on the rhythm of seasonal development of Prunella vulgaris L., a plant of the circumboreal region, have been available for the European part of Russia only. We examined the rhythm of seasonal development and minor life cycle of Prunella vulgaris L., the long-rhizome life form, in Siberia (Khakasia) in 2012-2013. Observations were carried out in a forest meadow every 5-7 days in the spring-summer-autumn period and 1-2 times a month in the winter. The development of dicyclic meso-rosetted and winter monocyclic semi-rosetted generative monocarpic shoots was observed using the method by I.G. Serebryakov. It was determined that the differentiation of vegetative and generative spheres in Prunella vulgaris L. shoots in Siberia occurs in early spring within the year of flowering. The species forms spring and autumn leaf generations and phenologically can be described as a summer-winter green plant with a long growing season. The period of shoot development from its initiation to dying the above-ground parts lasts 26 and 14 months in dicyclic meso-rosetted shoots and winter monocyclic semi-rosetted shoots, respectively. In both cases the development of the primordial shoot inside the bud lasts 8 months while the duration of shoot development after emerging from the bud varies. The latter phase lasts 18 months in dicyclic meso-rosetted shoots and only 6 months in winter monocyclic semi-rosetted shoots. Start times and duration of phenological phases (budding, flowering, fruiting and dissemination) are related to the weather conditions in the habitats including fluctuations in air temperature, precipitation patterns and types
Biological and structural peculiarities of Erodium tataricum Willd. cenopopulations in natural surroundings of Khakasia are examined. It is found out that the species make two vegetal forms under the changing environmental conditions. Ontogeny is complete, regeneration does not take place. Two types of ontogeny are found out: morphological and dynamic (according to the rate of growth). Examined Erodium tataricum Willd. cenopopulations are normal and incomplete. They are characterized by left-sided and bimodal ontogenetic spectra.
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