In freshwater environments, high biodiversity is commonly associated with habitat heterogeneity. River bends and meanders are particularly complex morphodynamic elements of watercourses. However, the specific spatio-temporal interactions between hydromorphology and the resident biota have scarcely been studied. This article reviews the relationships between hydraulic processes, and morphological units that are typical for meanders, and analyzes the concomitant spatial and temporal dynamics of habitats suitable for aquatic invertebrates. Flow in river bends is characterized by significant crossstream velocities, which modify primary flow patterns, and create helical flow trajectories. Consequently, boundary shear stresses at the river-bed are altered, so that complex erosion, transport, and accumulation processes characteristically shape bed and bank morphology. The diversity of substrate types and complex bathymetry in meanders provide a large variety of habitat conditions for benthic invertebrates within a relatively small spatial domain, which are connected via hydraulic pathways. Periodic reversal of hydromorphological processes between low and high flow, and seasonal growth of aquatic macrophytes creates spatio-temporal dynamics at the meso-and microhabitat scales. Such habitat dynamics increases benthic invertebrate diversity to the extent it is consistent with spatio-temporal scales of invertebrate mobility and life cycle. Furthermore, the presence of flow refugia, and hydraulic dead zones in meanders is essential to sustain species richness. This study concludes that meanders are highly complex morphodynamic elements that exhibit several self-regulating principles supporting invertebrate diversity and resilience in fluvial ecosystems.
[1] Shallow lateral shear layers forming between flows with different velocities, though essential for mixing processes in natural streams, have been examined only in laboratory settings using smooth, fixed-bed channels. This paper reports the results of an experimental study of a shear layer in a straight reach of a natural river where the layer, in contrast to the two-dimensional structure observed in the laboratory, is highly three-dimensional. The study included pronounced transverse pressure gradients, which influenced shear layer structure compared to flume experiments. It also introduces an analysis that complements conventional theory on mixing layers. The lateral velocity gradient between the flows downstream from a splitter plate placed in the river, the principal controlling factor, was adjusted for three experimental runs to determine the influence of different gradients on shear-layer dynamics. In each run, detailed three-dimensional measurements of mean and turbulent characteristics were obtained at five cross sections downstream from the splitter plate. Although experimental results agreed with conventional mixing-layer theories with respect to turbulence, the dynamics of the shear layers were dominated by the mean lateral fluxes of momentum. After re-examining the governing equations, we developed a parabolic equation describing the shear layer evolution and several scaling relations for essential terms of the energy budget: mean and turbulent lateral fluxes of momentum, turbulent kinetic energy, and dissipation rates. The study also provides insight into the spectral dynamics of turbulence in the shear layer and clarifies previous observations reported for confluences in natural streams.
Interactions between vegetation, flow and sediment are a key ingredient for the development of vegetated islands in highly dynamic, fluvial alpine ecosystems such as the Tagliamento River, north-east Italy. There has been substantial research on factors influencing the establishment of vegetation and feedback mechanisms between vegetation, hydraulic, and geomorphological processes in such environments. This has yielded the development of conceptual models identifying different trajectories of vegetation and landform development from bare gravel to established floodplain forest. Nevertheless, some of the finer-scale processes underpinning such interactions are not well understood and parameterisation concepts that augment our knowledge from process understanding to quantified data and prediction models are not available until now. This paper identifies mechanisms and parameters of vegetationflow interaction at the individual scale that are reflected at a patch or even at the channel scale. These mechanisms are reviewed from a multi-disciplinary perspective and concepts and analogies are proposed that provide ideas to progress research towards the development of predictive vegetationflow models. Such models must incorporate both hydraulic and ecological components and this is demonstrated for a simplified force-bending model of Salicaceae seedlings. The development of such models demands advances in the individual disciplines of hydraulics, morphology, plant ecology and biomechanics, which offers many possibilities for multidisciplinary research between these disciplines.
The effects of aquatic macrophytes on flow and turbulence were studied in a tightly curving meander bend. Three field measurement campaigns were carried out within a one year period to capture effects of seasonal changes in macrophyte cover. They comprised three‐dimensional velocity measurements and mappings of vegetation cover and bathymetry. Flow accelerates and converges over the run into an axial pool in a jet‐like flow pattern bordered by outer and inner bank flow separation zones. The jet and widening of the cross‐section anticipate helical flow until the second half of the bend, where an asymmetric pool developed. Submerged vegetation at the riffles preserves the jet at much lower discharges during the summer period by concentrating high momentum fluid near the surface. Plants locally modify the velocity and stress patterns, reduce bed shear stresses, create zones of fine sediment accumulation and reinforce the bed and banks with roots and rhizomes. Plant patches colonising the banks and the point bar confine secondary flow cells laterally and affect shape and magnitude of the transverse flow profiles near their edges. The morphology of the bend was very stable over the observation period and neither bank erosion nor pool scouring occurred. However, fine sediments accumulate within vegetation patches and in the recirculation zones while the remaining open areas tend to erode slightly. With the decay of macrophytes in winter, sediment accumulations are mobilised again and the bathymetry levels, supporting cyclic models of morphologic change in vegetated bends. In the second part of the paper, semi‐empirical models for the three predominant flow types were tested and discussed; velocity and stress models of vegetated mixing layers and plane turbulent jets, and Rozovskii's model for the transverse flow in bends. Copyright © 2012 John Wiley & Sons, Ltd.
Summary 1. Ship‐induced waves can affect the physical characteristics of lake and river shorelines, and laboratory studies have shown effects on littoral invertebrates. Here, we explored whether these effects could be observed under field conditions along a natural lake shore affected by wave sequences (trains) produced by boats. 2. Individuals of five invertebrate species (Bithynia tentaculata, Calopteryx splendens, Dikerogammarus villosus, Gammarus roeselii, Laccophilus hyalinus) were exposed to waves with increasing shear stress in five habitats differing in structural complexity. 3. Detachment of invertebrates increased with increasing shear stress and was best modelled using sigmoid response curves. Habitat structural complexity mitigated the effects of shear stress, and detachment rate was influenced more by habitat type than by species. A threshold (90% of the individual invertebrates unaffected) stress level of 0.64 N m−2 was found for a structurally complex reed habitat, compared to 0.37 N m−2 for a simple sand habitat. 4. Shear stress associated with wave trains created by recreational boating at a distance of 35 m from the shore and at a speed of 11 km h−1 resulted in 45% detachment of littoral invertebrates. Decreasing the boat‐to‐shore distance to 20 m increased wave shear stress by 30% and invertebrate detachments up to 75%. 5. Disturbance of littoral habitats and invertebrate assemblages are widespread in inland waters used for recreational and/or commercial navigation. Our findings show that the integrity of littoral zones of navigable surface waters could be much improved by implementing management measures such as physically protecting complex habitats with dense reed belts and tree roots, and reducing boat speeds and increasing their minimum shoreline distance.
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