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The response of wheat plants to different osmotic stress levels varied among the different organs root, shoot and spike and the situation of these organs with application of two Cu ++ levels 5 mM and 25 mM as CuSO 4 . The sensitivity of root organ was related with reduction in fresh, dry matter and length. This resulted from reduction of soluble sugar reflected a reduction in water uptake and K + content of the cell sap. In the moderate organ spike, the reduction in fresh, dry matter and length were concomitant with the accumulation of soluble sugar and a huge accumulation of soluble protein. In the higher organ shoot, this related with more water uptake which in turn induced an accumulation of soluble protein and cofactor K + content. It can be recorded that shoot was higher Na + accumulation than root and spike. Data also showed further stimulatory effect on growth parameters by Cu ++ applications with either concentration (7.5 mM or 25 mM). Irrigating the soil with either 7.5 or 25 mM CuSO 4 induced a huge accumulation in soluble sugar, soluble protein and nitrate reductase. Cupper treatment with either concentration 7.5 mM or 25 mM induced a marked decrease in Na + content at all OSL and has no significant change in the accumulation of K + in both shoot and spike whereas induced a huge accumulation in root organ. The synthesis of protein bands with molecular weight 32.3 KDa at −1.5 MPa NaCl level treated with either 7.5 mM or 25 mM Cu ++ concentrations was induced. Also the appearance of protein band with molecular weight 37KDa induced only at Cu ++ treatments with 25 mM concentration in both control and under different osmotic levels (0.0, −0.3 MPa, −0.9 MPa, −1.5 MPa NaCl).

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This investigation was conducting to explain that four barley genotypes varied in their drought tolerance according to their genotype and their tested organs. It can be recorded that growth parameters (fresh, dry matter and length, water content, leaf area and pigment contents) were decreased as decreasing M. C. in the soil. This indicated that Giza 123 was the superior in its drought tolerance and Giza 129 was the interior and both cv. Giza 2000 and cv. Giza 124 were the intermediated between them. This concomitant with increase in soluble sugar and soluble protein content of both organs in Giza 123 and shoot Ca ++ , this related with lower value of OP other than genotypes, in Giza 2000 this was related with a huge accumulation in soluble protein of shoot and root, shoot amino acids and root proline reach 3-folds than control plants as decreasing M. C. Whereas drought stress increased soluble protein only in Giza 124 while in Giza 129 decreasing M. C. increased soluble protein, amino acids and proline contents in shoot and root and shoot Ca ++. The values of OP increased as decreasing M. C. in four barley cultivars concomitant with their drought tolerance. Also, SA application was markedly enhanced the production of growth parameters in shoot and root with varied degree according to each tested barley genotypes. SA application was significantly increased OP in shoot, root and spike of barley pants. Spraying vegetative parts with 0.5 mM SA was markedly increased the soluble sugar, soluble protein and amino acids in shoot, root and spike of four barley cultivars. On the other side, SA application lowered the accumulation of proline in shoot and root of barley genotypes. SA treatment induced no significant change in K + , Ca ++ , and Mg ++ in shoot, root and spike of Giza 123, it significantly increased K + , Ca ++ , and Mg ++ in shoot and root of Giza 2000. SA application enhanced accumulation of K + , Ca ++ in shoot and root of Giza 124 and K + , Ca ++ and Mg ++ in three organs of Giza 129.

The present work was carried out to investigate the role of mycorrhiza inoculation at two harvesting stages (90-days and 30-days) of maize plants grown in pot experiment with different moisture content levels 100%, 70%, 50% and 20%. Drought stress tolerant in maize plant was varied in different organs of the same plants and also varied among different stage of plant development. The sensitivity of maize plants was related with reduction of root soluble sugar, shoot and root soluble protein at 30-days of plant harvesting, and soluble sugar and soluble protein in both organs of both harvesting stages. This related with reduction in OP and lowering of water uptake which induced a marked decrease in fresh and dry matter production in shoot and root of both harvesting stages. AM inoculation increase maize tolerant to drought stress presented in increasing growth parameters, chemical constituents and minerals contents compared with untreated plants. Proline content with AM inoculation was more or less unchanged in shoot of plant harvesting at 30-days and in root of plant harvesting at 90-days. However, a marked increase was induced in plant harvesting at 30-days and in shoot of plant harvesting at 90-days. Mycorrhiza inoculation induced a significant increase in OP value either compared with corresponding level or compared with control value 100% as in plant 30-days of harvesting or compared with control only as in plant harvesting after 90-days. AM infection with different moisture content levels measured by N-acetyl glucosamine content were not affected by drought stress. Results showed also that control roots contained N-acetyl glucosamine would be attributed to mycorrhiza and other fungi naturally present in soil.