Twelve localities of Lower Carboniferous strata in Scotland (Loch Humphrey Burn, Glenarbuck, Pettycur, Oxroad Bay and the Berwickshire localities of Cove, Burnmouth, Gavinton, Edrom, Foulden) and in France (Esnost, Roannais, Montagne Noire) have been investigated with particular regard to their anatomically preserved floras. New data on the composition and preservation of the assemblages, their geological setting and stratigraphical age, using palynology in particular, are presented. Present data suggest that four successive groups of floras obtained from these localities can be recognised: from the Montagne Noire (mid to late Tournaisian), from the Berwickshire localities and Oxroad Bay (late Tournaisian to early Viséan), from Loch Humphrey Burn and possibly Glenarbuck (mid Viséan) and from Roannais, Esnost, and Pettycur (late Viséan). The similarities and differences between the floras are discussed with particular emphasis on stratigraphical as opposed to ecological controls. Comparisons are made with the New Albany Shale floras of the U.S.A., the Saalfeld and Glätzish-Falkenberg floras from the German Democratic Republic. The data suggest that the ferns and pteridosperms show the most striking changes through the Lower Carboniferous strata.

SynopsisThis paper provides a comprehensive account of the distribution of miospore species which are considered to be stratigraphically significant in the Dinantian (Calciferous Sandstone Measures and Lower Limestone Group) of the Midland Valley of Scotland. Five miospore concurrent range zones are recognised in these beds:Tripartites vetustus—Rotaspora fracta (VF) ZoneRaistrickia nigra—Triquitrites marginatus (NM) ZonePerotrilites tessellatus—Schulzospora campyloptera (TC) ZoneLycospora pusilla (Pu) ZoneSchopfites claviger—Auroraspora macra (CM) ZoneThe distribution of these zones in the sections studied provides a method of correlation in a series of rocks, which are largely of non-marine origin and notable for their lack of stratigraphically useful macrofossils and for marked variations in facies and thickness. The zones have also been proved in sections of Lower Carboniferous rocks in the Northumbrian trough, thereby providing a potential method of correlation between the two areas.Two new genera, Colatisporites and Biannulatisphaerites, are described and one Tricidarisporites emended. Fifteen new species and one new type are described, three species recombined and three emended. Representative assemblages of the miospore zones are illustrated.

Reconstructions based on biogeography, palaeomagnetism and facies distributions indicate that, in later Palaeozoic time, there were no wide oceans separating the major continents. During the Silurian and Early Devonian time, many oceans became narrower so that only the less mobile animals and plants remained district. There were several continental collisions: the Tornquist Sea (between Baltica and Avalonia) closed in Late Ordovician time, the Iapetus Ocean (between Laurentia and the newly merged continents of Baltica and Avalonia) closed in Silurian time, and the Rheic Ocean (between Avalonia and Gondwana and the separate parts of the Armorican Terrane Assemblage) closed (at least partially) towards the end of Early Devonian time. Each of these closures was reflected by migrations of non-marine plants and animals as well as by contemporary deformation. New maps, based on palaeomagnetic and faunal data, indicate that Gondwana was close to Laurussia during the Devonian and Carboniferous periods, with fragments of Bohemia and other parts of the Armorican Terrane Assemblage interspersed between. It follows that, after Early Devonian time, the Variscan oceans of central Europe can never have been very wide. The tectonic evolution of Europe during Devonian and Carboniferous time was thus more comparable with the present-day Mediterranean Sea than with the Pacific Ocean.

The Central Irish Sea area, from Kish Bank to St. George's Channel and Cardigan Bay, consists of a series of Late Palaeozoic to Cenozoic extensional and transtensional basins which have experienced a multiphase inversion history. Potential hydrocarbon source rocks of Carboniferous and Jurassic ages have been recognised in this area. In the Kish Bank and Central Irish Sea Basins, maturation modelling of these source rocks is hampered by the severely truncated rock record and by the relative paucity of vitrinite throughout much of the preserved post‐Palaeozoic (Triassic) section. Vitrinite reflectance data from six exploration wells have been used to quantify the peak palaeotemperatures attained by the rocks in this area and to estimate the magnitude of net exhumation at these locations. An apparent palaeogeothermal gradient of ˜26°C/km is recorded by the Jurassic sediments in well 42/21‐1, whereas significantly higher palaeogeothermal gradients of 74–78°C/km are interpreted for the Westphalian/Stephanian sediments in the area. At least two periods of rock exhumation have occurred; during the Late Carboniferous‐Late Permian, and again sometime between the latest Jurassic and early Tertiary. Estimates of net exhumation vary from ˜350 m at well 42/12‐1 to ˜1,900 m at well 42/17–1. Our interpretation suggests that the higher palaeogeothermal gradients recorded by the Westphalian/Stephanian sections reflect elevated heat flows during Stephanian to Early Permian times.

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