No abstract

The Foresta ichnosite is well known for preserving some of the oldest human fossil footprints recorded in Europe so far. This research aims to: i) describe new footprints that are larger than those already reported, some of which form a new trackway that moves in the opposite direction to all the others; ii) announce the discovery of some stone tools also in the surroundings of the Foresta ichnosite. The new results increase the total number of human fossil footprints to at least 81, specify the direction and the number of footprints of Trackway C, and identify three new directions of walking at the site. More compelling and complete estimates of the dimensional range of all ichnological evidence enables us, furthermore, to estimate the number of trackmakers walking on the trampled surface as a minimum of five, one of them likely being an adult male. The general shape of all the recorded footprints suggests that the Foresta trackmakers share some similarities with those at Sima de los Huesos, and belong to the same taxonomical group as the Ceprano skull. All the new evidence enables us to better understand the presence of hominin populations in the Roccamonfina volcano area during the Middle Pleistocene. © 2020 John Wiley & Sons, Ltd.