Imagoes, subimagoes and eggs of Indobaetis costai Müller-Liebenau & Morihara 1982 are described for the first time; additional larval characters of I. costai and I. starmuehlneri Müller-Liebenau & Morihara 1982 are reported. A new species Indobaetis microfolius sp.n. is described based on larvae, a male subimago reared from larva and a female subimago extracted from a mature larva. Baetis michaelohubbardi (Selva-kumar et al. 2012) comb.n. is proposed for a species originally attributed to Indobaetis, based on the presence of hind protoptera, three maxillary dentisetae and other characteristics. Newly discovered diversity within the holophyletic taxon Indobaetis disproves a formerly erected idea about the close relationship of Indobaetis and the muticus-group. Thus, the senior genus-group name for the holophylum including the muticus-group is Takobia/g1; in rank-based nomenclature the names Alainites Waltz & McCafferty (in Waltz et al.) 1994 and Acerbaetis Kang & Yang (in Kang et al.) 1994 should be junior synonyms of Takobia Novikova & Kluge 1987 (new synonymy). The Neotropical taxon Caribaetis Kluge 1992 is accepted as genus (stat.n.), rather than a synonym of Fallceon, based on the original diagnosis of the subgenus Caribaetis and taking into account the significance of such character as presence of a posterior seta on the claw, shared with Indobaetis, Baetodes and some other taxa; resultant new combinations are proposed: Caribaetis planifrons (Kluge 1992) comb.n., Caribaetis alcarrazae (Kluge 1992) comb.n. and Caribaetis grandis (Gonzalez-Lazo & Salles 2007) comb.n. A new species synonymy in Fallceon is proposed, based on the presence of a hind wing costal hook in both species: Fallceon poeyi (Eaton 1995) = F. nikitai McCafferty & Lugo-Ortiz 1994 syn.n.
includes 2 species inexpectatum [Pseudocloeon] and gnom [Pseudocloeon]; their larvae, subimagoes, imagoes and eggs are described. Group sibiricum includes 3 species sibiricum [Pseudocloeon], diptera sp.n. [Acentrella] (new species from Siberia and Russian Far East) and scabriventris sp.n. [Acentrella] (new species from Kyrgyzstan and Tajikistan); their larvae, subimagoes, imagoes and eggs are described. African and Oriental representatives of Acentrella/fg1 constitute a holophyletic taxon Jubabaetis/g1 characterized by an autapomorphy in structure of left mola. Jubabaetis/g1 is divided to African taxon Tanzaniops/g(1) and Oriental taxon Jubabaetis/g2. A new species lunamontana sp.n. [Acentrella (Tanzaniops)] is described as larvae, subimagoes and imagoes from Uganda. Holophyletic taxon Jubabaetis/g2 is divided into a plesiomorphon Liebebiella/g(1) and holophyletic taxa Jubabaetis/g3 and Platybaetis/g(1). Three species of Liebebiella/g(1) are described as larvae, subimagoes and imagoes: verum [Pseudocloeon] (= deigma [Liebebiella], syn.n.), bispinosa sp.n. [Acentrella (Liebebiella)] and cylindroculata sp.n. [Acentrella (Liebebiella)] (new species from Sulawesi). Two species of Platybaetis/g(1) from Sulawesi are described as larvae, subimagoes and imagoes: mamasae sp.n. [Platybaetis] and wallacei [Platybaetis]. (Tanzaniops)]. Ãîëîôèëåòè÷åñêèé òàêñîí Jubabaetis/ g2 äåëèòñÿ íà ïëåçèîìîðôîí Liebebiella/g(1) è ãîëî-ôèëåòè÷åñêèå òàêñîíû Jubabaetis/g3 è Platybaetis/g(1). Ïî ëè÷èíêàì, ñóáèìàãî è èìàãî îïèñûâàþòñÿ òðè íîâûõ âèäà Liebebiella/g(1): verum [Pseudocloeon] (= deigma [Liebebiella], syn.n.), bispinosa sp.n. [Acentrella (Liebebiella)] è cylindroculata sp.n. [Acentrella (Liebebiella)] (íîâûå âèäû èç Ñóëàâåñè). Ïî ëè÷èíêàì, ñóáèìàãî è èìàãî èç Ñóëàâåñè îïèñûâàþòñÿ äâà âèäà Platybaetis/g(1): mamasae sp.n. [Platybaetis] è wallacei [Platybaetis]. ! Systematics of the mayfly taxon Acentrella, with description of new species generic name Pseudocloeon, this species is regarded as the type species of the generic name Pseudocloeon by monotypy. As it was discovered later, loss of hind wings took place independently in many non-related taxa within Baetovectata [Novikova & Kluge, 1987; Waltz & Mc-Cafferty, 1987ab]; so the genus Pseudocloeon, as characterized and outlined by Klapalek, is polyphyletic. Recently this polyphyletic group is split into a number of smaller, more natural taxa. The question, to which of these taxa the old name Pseudocloeon should be applied, remains to be opened. The type species Pseudocloeon kraepelini was described basing on specimens, collected by Karl Kraepelin 24.II12.III.1904 in Buitenzorg (now Bogor) in Java. Details of this trip can be learned from the paper by Weidner [1972]. After this, nobody vas able to collect this species. The type material is deposited in Zoologisches Museum in Hamburg. Lectotype was designated and redescribed by Waltz and McCafferty in 1985; its photos, made by Dr Ralph Peters in 2009, are available from web
A new tribe Labiobaetini tribus n. is established for a plesiomorphon Labiobaetis Novikova & Kluge 1987 and a holophyletic taxon Pseudopannota Waltz & McCafferty 1987. New synonymy Pseudopannota Waltz & McCafferty 1987 = Ophelmatostoma Waltz & McCafferty 1987, syn. n. is proposed. A new species Pseudopannota pannota sp. n. closely related to P. berthrandi (Demoulin 1967) is described from Zambia based on imagoes of both sexes reared from larvae. Pseudopannota camerunense (Ulmer 1920) comb. n. is redescribed from Zambia based on imagoes of both sexes reared from larvae. A new species Pseudopannota fusca sp. n. is described from Uganda based on larvae and subimagoes of both sexes extracted from mature larvae. Pseudopannota maculosa (Crass 1947) is reported from Zambia based on larvae. Labiobaetis tenuicrinitus (Kopelke 1980) comb. n. (= Baetis spatulatus Gillies 1994 syn. n.) is redescribed from Uganda based on imagoes of both sexes reared from larvae.
Two new species, Anafroptilum orthostylus sp. n. and Anafroptilum odontostylus sp. n. are described from Thailand based on imagoes reared from larvae. Both new species lack hind wings; larvae of both species have an unusual prominent spine near cercus apex; other characters, especially shape of male imaginal gonostyli and shape of larval tergalii, are quite diverse in these two species. The genus Anafroptilum, hitherto known from the East Palaearctic [as a single species A. kazlauskasi (Kluge 1983)] and from Nearctic [as eight species reported by Jacobus & Wiersema (2014)], is reported from Oriental Region for the first time.
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