The fossil record provides a fantastic laboratory to explore the effects of past climatic and environmental changes on biodiversity and morphology (Barnosky et al., 2003;Blois & Hadly, 2009;Dietl & Flessa, 2011). Rodents are of particular interest because of their high taxonomic diversity during the Cenozoic (Samuels & Hopkins, 2017); their small body size, short breeding cycles, and ecological adaptations make them helpful indicators of environmental change (Chaline, 1977). One group of rodents that is expected to closely track the environment through time is fossorial rodents (e.g., Calede et al., 2011); a group of animals whose distribution and abundance is dependent on soil and environmental characteristics (e.g., Lazo-
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