N the first article of this series, GEISSMAN and MEHLQUIST (1947) described the I chemical effects that accompanied alterations in the genetic constitution of carnations with respect to the factors S, R and A&. It was found that only R affected the degree of nuclear hydroxylation of the anthocyanin pigments of the flowers. Cyanic forms that contained R in homozygous or heterozygous condition contained cyanidin glycosides; the homozygous recessives (rr) contained pelargonidin derivatives. At the time of that study the number of genetically analyzed non-cyanic forms was limited, and of the whites and yellows only rr specimens were a t hand. It was found that these white and yellow rr flowers (which contain no anthocyanins)
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