A cellular automata computer model for the interactions between two plant species of different growth forms-the lime hairgrass Deschampsia caespitosa (L.) P. Beauv., a sod cereal, and the moneywort Lysimachia nummularia L., a ground creeping perennial herb-is considered. Computer experiments on the self maintenance of the populations of each species against the background of a gradual increase in the share of randomly eliminated individuals, coexistence of the populations of two species, and the effect of the phy togenous field have been conducted. As has been shown, all the studied factors determine the number of indi viduals and self sustainability of the simulated populations by the degree of their impact. The limits of action have been determined for individual factors; within these limits, the specific features in plant reproduction and dispersal provide sustainable coexistence of the simulated populations. It has been demonstrated that the constructed model allows for studying the long term developmental dynamics of the plants belonging to the selected growth forms.
The proposed model is the version of well-known biomass production model 3-PG (Physiological Principles Predicting Growth), which allows for calculation of biomass production in
dependence of consumed soil nitrogen and available solar radiation. The model utilizes the concept of modifiers, i.e. functions describing the effect of tree age and environmental
factors (air temperature and humidity, soil moisture, carbon dioxide concentration) on productivity. To make the model applicable to mixed forests of European Russia, the substantial
modifications were implemented. In particular, more detailed response functions to air temperature, soil moisture and absorbed nitrogen were introduced. We also implemented new procedure
of calculation of light use efficiency taking into account the difference between shade-tolerant and shade-intolerant tree species. The rank distribution equation was used for the
description of an increment allocation to different tree biomass compartments. Model parameters were estimated for the 12 most common tree species of European Russia. The model
was implemented as sub-routine for calculation of biomass production in forest ecosystem model EFIMOD 2. The model performance was tested against the wide range of environmental conditions.
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