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The tyrannosauroid fossil record is mainly restricted to Cretaceous sediments of Laurasia, although some very fragmentary Jurassic specimens have been referred to this group. Here we report a new basal tyrannosauroid, Guanlong wucaii gen. et sp. nov., from the lower Upper Jurassic of the Junggar Basin, northwestern China. G. wucaii is the oldest known tyrannosauroid and shows several unexpectedly primitive pelvic features. Nevertheless, the limbs of G. wucaii share several features with derived coelurosaurs, and it possesses features shared by other coelurosaurian clades. This unusual combination of character states provides an insight into the poorly known early radiation of the Coelurosauria. Notably, the presumed predatory Guanlong has a large, fragile and highly pneumatic cranial crest that is among the most elaborate known in any non-avian dinosaur and could be comparable to some classical exaggerated ornamental traits among vertebrates.

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The lithostratigraphic interval between the Taber and Lethbridge coal zones in the upper portion of the nonmarine Judith River Group of southeastern Alberta is divisible into two lithostratigraphic units separated by a regionally extensive and diachronous discontinuity. The lower unit, referred to here as the Oldman Formation, is characterized by very fine grained to fine-grained sandstones that contain fewer than 2% volcanic rock fragments; sandstone bodies with numerous sets of horizontally stratified sandstone, showing little or no evidence of lateral accretion; siliceous paleosols (ganisters); and a relatively high gamma-ray signal in the upper half of the formation. The Oldman Formation comprises deposits of a low-sinuosity, perhaps ephemeral fluvial system that originated in the southern Cordillera of Canada and northern Montana and flowed northeastward, perpendicular to the axis of the Alberta Basin.The upper unit is assigned to a new formation, the Dinosaur Park Formation, and is characterized by fine- to medium-grained sandstones with up to 10% volcanic rock fragments; sandstone bodies that exhibit lateral-accretion surfaces in the form of inclined heterolithic stratification; numerous articulated dinosaurs and dinosaur bone beds; and a relatively low gamma-ray signal in the lower half of the formation. The Dinosaur Park Formation comprises deposits of a high-sinuosity, fluvial-to-estuarine system that originated in the north and central Cordillera and flowed southeastward, subparallel to the axis of the Alberta Basin.40Ar/39Ar and K–Ar dating of Judith River Group bentonites shows that the contact between the Oldman and Dinosaur Park formations becomes younger toward the south and southeast. These data also demonstrate that the Dinosaur Park Formation clastics migrated southeastward at a rate of approximately 130–140 km/Ma, gradually overstepping the Oldman Formation elastics.The widely recognized north-to-south increase in intensity of overthrust loading along the western margin of the Alberta Basin during the Late Cretaceous is thought to be responsible for (i) differences in accommodation space for the proximal portions of the Oldman and Dinosaur Park formations, and (ii) the establishment of a southerly tilt in the Alberta Basin leading to the southeastward migration of the Dinosaur Park Formation elastics. In the northern portion of the basin, relatively lower rates of subsidence, combined with periods of isostatic rebound in the foredeep, resulted in the southeastward migration of Dinosaur Park Formation elastics as sediment input exceeded accommodation space. In the southern portion of the basin, relatively higher rates of subsidence and little isostatic rebound acted to trap coarse-grained Oldman Formation elastics in the foredeep and may have led to periods of sediment starvation in more distal portions of the basin. An inferred lower depositional slope associated with the Dinosaur Park Formation (relative to the Oldman Formation) is thought to have resulted from gradual loading of the basin as Dinosaur Park Formation elastics migrated southeastward or some form of tectonically induced subsidence.

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Multi-kilometre-thick Jurassic–Cretaceous-age sedimentary successions exposed in the southern, northwestern, and northeastern regions of the Junggar Basin display a consistent and correlative stratigraphy comprising four, stacked second-order megasequences: Badaowan, Sangonghe, Shishugou, and Kalaza. Each consists of a basal erosional unconformity or discontinuity surface and lower, middle, and upper units that are interpreted as forestepping, backstepping, and aggradational systems tracts, respectively. Each megasequence is interpreted as recording an upsection shift from active tectonism and uplift to tectonic quiescence with associated changes in crustal response and sediment supply. Basin-wide analysis of megasequences indicates that tectonism was intermittent and regionally variable. A maximum phase of subsidence and sediment accommodation is recorded in the middle unit of each megasequence and correlates with a notable abundance of fossil vertebrates suggesting a primary tectonic and basin-response control on fossil preservation. Seasonally dry climatic conditions were developed first in the northeastern region of the basin during the Pliensbachian, followed by basin-wide seasonal dryness during the Bajocian. Seasonally dry climatic conditions were permanently established across the basin by the Oxfordian and intensified during the Early Cretaceous. A seasonally dry climate from Oxfordian through the Early Cretaceous correlates positively with the widespread presence of fossil vertebrates and suggests an additional climatic control on fossil preservation.

Because the Bayan Mandahu redbeds of Inner Mongolia share similar sedimentary facies and fossil assemblages with the Djadokhta Formation of pre-Altai Gobi, the two units are interpreted as stratigraphic correlatives, both of Campanian age. Sedimentary facies indicate that the Bayan Mandahu redbeds were deposited in semiarid, alluvial to eolian environments. An assemblage of fossil vertebrates found in the Bayan Manduhu consists of ceratopsian, ankylosaurian, and theropod dinosaurs; turtles; crocodiles; and small lizards and mammals. Six different kinds of fossil vertebrate eggs are present. The most common fossil vertebrates occur in association with eolian deposits and are interpreted as the remains of autochthonous "faunal" components, many of which died in situ during sandstorm events. In contrast, rare and fragmentary specimens of large dinosaurs occur in coarse-grained alluvial deposits and are interpreted as the remains of allochthonous faunal components. The low diversity of this fossil assemblage and overall small to medium size of its constituents indicate a relatively stressed paleoenvironment, an interpretation which is compatible with our sedimentological conclusions. A diverse trace fossil assemblage is present and includes rhizoliths and endogenic traces. Endogenic traces are well preserved and typically associated with eolian deposits, suggesting that the deposits were at least seasonally damp and cohesive. The opinion that the Late Cretaceous Gobi Basin was a large inland lake, still advocated by some authors, cannot be maintained within the context of our sedimentologic and paleontologic data. In contrast with the perennial lacustrine sedimentation that was characteristic of the underlying Lower to lower Upper Cretaceous units in the Gobi Basin, the Bayan Mandahu redbeds and correlative Djadokhta Formation mark a pronounced shift toward eolian and intermittent lacustrine sedimentation in an increasingly arid climate.

A high-resolution biostratigraphic analysis of 287 dinosaurian macrofossils and 138 bonebeds in the Edmonton Group (Upper Cretaceous) of southern Alberta provides evidence for at least three dinosaurian assemblage zones in the Horseshoe Canyon Formation (HCFm). From bottom to top the zones comprise unique assemblages of ornithischians and are named as follows: (1) Edmontosaurus regalis – Pachyrhinosaurus canadensis (lower zone); (2) Hypacrosaurus altispinus – Saurolophus osborni (middle zone); and (3) Eotriceratops xerinsularis (upper zone). Whereas the lower and middle zones are well defined and based on abundant specimens, the validity of the uppermost zone (E. xerinsularis) is tentative because it is based on a single specimen and the absence of dinosaur taxa from lower in section. The transition from the lower to the middle zone coincides with the replacement of a warm-and-wet saturated deltaic setting by a cooler, coastal-plain landscape, characterized by seasonal rainfall and better-drained substrates. Whereas changes in rainfall and substrate drainage appear to have influenced the faunal change, changes in mean annual temperature and proximity to shoreline appear to have had little influence on faunal change. We speculate that the faunal change between the middle and upper zones also resulted from a change in climate, with ornithischian dinosaurs responding to the re-establishment of wetter-and-warmer climates and poorly-drained substrates. Compared with the shorter-duration and climatically-consistent dinosaurian assemblage zones in the older Dinosaur Park Formation of southern Alberta, HCFm assemblage zones record long-term morphological stasis in dinosaurs. Furthermore, the coincidence of faunal and paleoenvironmental changes in the HCFm suggest climate-change-driven dinosaur migrations into and out of the region.