Changes in the weed seedbank due to crop production practices are an important determinant of subsequent weed problems. Research was conducted to evaluate effects of primary tillage (moldboard plowing and chisel plowing), secondary tillage (row cultivation), and herbicides on weed species changes in the soil seedbank in three irrigated row crop rotational sequences over a 3-yr period. The cropping sequences consisted of continuous corn for 3 yr, continuous pinto beans for 3 yr, or sugarbeets for 2 yr followed by corn in the third year. Cropping sequence was the most dominant factor influencing species composition in the seedbank. This was partly due to herbicide use in each cropping sequence producing a shift in the weed seedbank in favor of species less susceptible to applied herbicides. A comparison between moldboard and chisel plowing indicated that weed seed of predominant species were more prevalent near the soil surface after chisel plowing. The number of predominant annual weed seed over the 3-yr period increased more rapidly in the seedbank after chisel plowing compared to moldboard plowing unless effective weed control could be maintained to produce a decline in seedbank number. In this case, seedbank decline was generally more rapid after moldboard plowing. Row cultivation generally reduced seedbanks of most species compared to uncultivated plots in the pinto bean and sugarbeet sequences. A simple model was developed to validate the observation that rate of change in the weed seedbank is influenced by type of tillage and weed control effectiveness.

Summary. Post harvest soil samples taken during the autumn of 1985 and 1986 were split and estimates made of the weed seedbank using two methods: (1) a physical separation of seed from the soil mineral fraction by a sieving/flotation procedure and (2) by placing soil in shallow containers in a greenhouse where seeds could germinate and be periodically counted over a period of eight months. Seedbank estimates derived from each procedure were analysed to determine the suitability of each technique for detecting treatment differences from an experiment evaluating tillage/herbicide effects on weed populations. Both techniques were suitable for determination of seedbank changes due to different tillage treatments and herbicide inputs. The two techniques also proved effective for detection of individual species in the seedbank and the two techniques provided comparable estimates of the relative density of individual weed species in the seedbank. Weed seedbank estimates obtained by the physical extraction procedure from the autumn 1985 soil samples were correlated with weed seedling counts made in the spring of 1986. In most cases, weed seedlings represented less than 10% of the number of seeds estimated by physical extraction the preceding autumn. Individual species seed estimates and subsequent weed counts were poorly correlated which indicated that the seed count estimates alone were poor predictors of weed flora.

Studies were conducted to determine the relative fitness and competitive ability of an acetolactate synthase (ALS) inhibitor–resistant (R) downy brome biotype compared with a susceptible (S) biotype. In previous research, the mechanism of resistance was determined to be an altered ALS enzyme. Seed germination of the R biotype was compared with that of the S biotype at 5, 15, and 25 C. There were no different germination characteristics between R and S biotypes at 15 and 25 C. However, the R biotype germinated 27 h earlier than the S biotype and had reached over 60% germination when the S biotype initially germinated at 5 C. Under noncompetitive greenhouse conditions, growth of the R biotype was similar to that of the S biotype on the basis of shoot dry weight, leaf area, and plant height. Seed production of the R biotype was 83%, when compared with the S biotype, but seeds of the R biotype were larger than those of the S biotype. Replacement series experiments were conducted in the greenhouse to determine the relative competitive ability of R and S biotypes. No difference in competitive ability was observed between R and S biotypes on the basis of shoot dry weight, leaf area, or plant height. Thus, it appears that ALS-resistance trait is not associated with growth penalty in either noncompetitive or competitive conditions. In the absence of ALS inhibitors, these results suggest that the R biotype would remain at a similar frequency in a population of R and S biotypes.

Research was conducted to evaluate the effects of primary tillage (moldboard plowing and chisel plowing), secondary tillage (row cultivation), and herbicides on weed species changes in the soil seed bank in three irrigated row cropping sequences over a 3-yr period. The cropping sequences consisted of continuous corn for 3 yr (CN), continuous pinto beans for 3 yr (PB), and sugarbeets for 2 yr followed by corn in the third year (SB). A comparison between moldboard and chisel plowing indicated that weed seed were more prevalent near the soil surface after chisel plowing. The density of certain annual weed seed over the 3-yr period increased more rapidly in the seed bank after chisel plowing compared to moldboard plowing. Species exhibiting the most pronounced increase included hairy nightshade and stinkgrass in the PB cropping sequence and redroot pigweed and common lambsquarters in the SB sequence. Conversely, kochia seed density in the SB sequence decreased more rapidly in chisel-plowed plots. Row cultivation generally reduced seed bank densities of most species compared to uncultivated plots. Herbicide use in each cropping sequence produced a shift in the weed seed bank in favor of species less susceptible to applied herbicides. In particular, seed of hairy nightshade became prevalent in the PB cropping sequence, and seed of kochia, redroot pigweed, and common lambsquarters became prevalent in the SB sequence.

Jointed goatgrass (Aegilops cylindrica) is a serious problem for winter wheat producers throughout the western U.S. Interference from this weed can severely reduce grain yield and contaminate harvested grain, resulting in dockage losses. There are currently no selective herbicides registered for controlling jointed goatgrass in wheat. Imazamox, an imidazolinone herbicide, was applied to an imidazolinone herbicide-resistant (IMI) wheat mutant of the winter wheat cultivar ‘Fidel.’ Jointed goatgrass control from spring postemergence application of imazamox ranged from 61 to 97% when applied at 36 g/ha. Negligible crop injury from imazamox treatment was observed at 36 g/ha at several locations under dryland environments in the U.S. Pacific Northwest. Wheat yield was increased 19 to 41% by imazamox treatment in three of four experiments. Percent dockage resulting from jointed goatgrass spikelet contamination of harvested wheat grain was eliminated by imazamox treatment. Introduction of the IMI trait into commercial wheat cultivars could provide an effective method for selective control of jointed goatgrass in winter wheat.

Control of rattail fescue, a winter annual grass, can be difficult in spring or winter wheat. Although rattail fescue is not a new weed species in the Pacific Northwest, occurrences have been increasing in circumstances where soil disturbances are minimized, such as in direct-seed cropping systems. To develop integrated management strategies for rattail fescue, information is needed on the longevity of seed viability in the soil, the presence of seed dormancy, vernalization requirements, and optimal environmental conditions for seed germination and establishment under field conditions. Controlled experiments on the biology of rattail fescue indicated that newly mature seed required an afterripening period of 1 to 12 mo to obtain high levels of seed germination, depending on germination temperature. Maximum seed germination was observed at constant day/night temperatures of approximately 20 C from thermogradient plate studies. Germination tests from seed burial studies indicated that a majority of buried seed was not viable after 2 to 3 yr. Field-grown rattail fescue plants required vernalization to produce panicles and germinable seed. A short afterripening period, cool germination temperature, and vernalization requirements support the classification of rattail fescue as a winter annual. This information will facilitate development of rattail fescue management systems, including crop rotations and various control tactics such as tillage or herbicide application timing during fallow periods.

Above-ground seedling development was characterized for five annual grass weeds: downy brome, bulbous bluegrass, jointed goatgrass, Italian ryegrass, wild oat; and three cereals: winter wheat, winter barley, and winter triticale in field experiments over two years. The rate of leaf production on the main stem of each species was linearly related to cumulative growing degree days (GDD) since planting. Leaf production rates were faster for bulbous bluegrass, downy brome, Italian ryegrass, wild oat, and barley than for wheat, triticale, and jointed goatgrass. The main stem development stage when individual tillers appeared was similar in all species except under poor seedbed conditions in 1991, in which case lower-node tillers were delayed in the cereals and jointed goatgrass, but not in most of the weed species. Bulbous bluegrass, downy brome, and barley had the same percentage of plants produce the first four primary tillers on the main stem in both years; the other species showed more year-to-year variation. Seedling heights at full emergence were generally greater for large-seeded species. Small-seeded species compared to large-seeded species tended to have greater relative increases in plant height over time. Knowledge of comparative development rates between these weeds and cereals could provide information for development of growth models for each of the species and could also improve understanding of the competitive relationships between grass weeds and cereal grains.

Field experiments were conducted to determine the effect of different rates and application times of MON 37500 on downy brome control in winter wheat and on rotational crop injury. In southeastern Washington, MON 37500 applied postemergence at 36 to 72 g ai/ha in the fall and spring controlled downy brome 90 to 97% and 45 to 71%, respectively. MON 37500 was applied postemergence to winter wheat at 18, 36, and 72 g/ha during fall 1995 and spring 1996 near Moscow, ID; Pendleton, OR; and Endicott, WA, to determine its effect on rotational crops planted during spring 1997. At Pendleton, OR, fall applications of MON 37500 at 18, 36, and 72 g/ha applied to actively growing wheat reduced seed yield of spring barley and pea 13 to 59% and 26 to 29%, respectively, when these crops followed winter wheat in the rotation. At this site, spring applications of 36 and 72 g/ha reduced seed yield of spring barley 17 to 34%; when applied in the fall or spring, MON 37500 reduced canola yield 31 to 73%. MON 37500 at 18 g/ha in the spring reduced pea biomass 32% at Endicott. Aboveground biomass and seed yield of barley, pea, lentil, and mustard at Moscow were not affected by MON 37500 carryover. However, using a grain sorghum bioassay, predicted final concentrations of MON 37500 in soil at all three locations 360 d after treatment (DAT) was similar, ranging from 0.5 to 2 ng ai/g. In growth chamber herbicide dose–response studies, grain sorghum shoot height was reduced more at lower doses (0.025 to 5 ng/g) of MON 37500 in Pendleton soil than in soils from the other field locations. Perhaps the greater rotational crop injury observed at the Pendleton location was partly related to greater soil activity of the herbicide.