Food resource is one of the most crucial exogenous factors for determining species distribution and abundance, as well as the ecological structure of populations and communities (Stergiou & Karpouzi, 2002;Wootton, 2012). The feeding activity of animals provides energy for the maintenance of life, somatic growth and reproduction, and forms a link between different trophic levels, thus inducing food competition. Producers and consumers are linked by their feeding activities through intra-and interspecific competition
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