S U M M A R YSix plant species were compared as food for sheep: chickweed (Stellaria media (L.) Vill.), dandelion (Taraxacum officinale Weber), dock (Rumex obtusifolius L.), ribwort (Plantago lanceolata L.), spurrey (Spergula arvensis L.) and perennial ryegrass (Lolium perenne L.). In two experiments at Aberystwyth in 1985 and 1986, the species were fed to lambs as the total diet, after artificial drying. In another experiment in 1987, they were fed fresh, in 1-5 minute meals, to yearling sheep.The dicotyledonous species had a lower proportion of cell wall (neutral detergent fibre) in dry matter than perennial ryegrass. The true dry matter digestibility of chickweed, dandelion, dock and leafy ribwort was as high as that of ryegrass, but the in vivo digestibility of all five dicotyledonous species was lower than that of ryegrass. The voluntary intake of the dicotyledonous species was higher than might have been predicted from their digestibility. The voluntary intake of spurrey was > 20 % higher than that of ryegrass.When the species were fed fresh, spurrey had the highest rate of intake as a consequence of a high chewing rate and a low requirement for number of chews/g of dry matter consumed. Chewing rate was higher on all the dicotyledonous diets than on ryegrass. The rate of intake of dandelion was above average, but that of dock was low and variable, particularly when the plants had been chopped.

SUMMARYIn a field which had been grazed by dairy cows for 26 years, the bulk density of both total soil and fine earth increased rapidly with depth down to 10–12 cm, but decreased below this depth. The bulk density was approximately twice as great in the 10–12 cm as in the 2–4 cm layer. Total porosity in the 10–12 cm layer was only 22%.Slitting the soil to penetrate the compacted layer approximately doubled net herbage accumulation and the net uptake of N, P and K, raising herbage production and uptake of nutrients from a low level to an acceptable one. Slitting tended to increase the concentrations of nitrate-N and K in herbage, but had little effect on the concentration of total N. Slitting increased the weight of ash-free root in the 10–20 cm depth range.

SUMMARYFifteen grassland species were grown in a heated glasshouse in February-April 1985. The harvested herbage was separated into leaf and ‘stem’ and analysed for neutral detergent fibre, digestibility, physical breakdown when macerated, fibrosity, water soluble carbohydrate and N, P, K, Ca, Mg and Na.Lolium perenne was digestible, but rather high in neutral detergent fibre and not readily broken down by maceration. The other grasses tested, Holcus lanatus, Poa annua and Glyceria fluitans, tended to be less digestible and higher in neutral detergent fibre than L. perenne.The leaves of all 11 dicotyledonous species were much lower in neutral detergent fibre than the leaves of the grasses, and most broke down readily when macerated. The stems of Medicago saliva and of two shrub species (Lonicera periclymenum and Prunus spinosa) were the highest in neutral detergent fibre.Rumex obtusifolius, Spergula arvensis and Stellaria media were high in Mg; Plantago lanceolata, S. arvensis and S. media were high in Na; and S. arvensis was high in P.Three perennial and two annual herbaceous, dicotyledonous non-legumes were selected as being of sufficient promise and interest to be compared with Lolium perenne in feeding experiments: Plantago lanceolata, Taraxacum officinale, Rumex obtusifolius, Spergula arvensis and Stellaria media.

Six varieties of white clover, each grown with perennial ryegrass, four intervals between cuts and two levels of applied nitrogen in all combinations, were compared in a field experiment during the first 27 months after sowing.Increasing the interval between harvests from 3 or 4 to 8-12 weeks increased the yield of white clover and generally did not reduce the proportion of clover in total herbage. Increasing the interval between harvests reduced the number of grass tillers but increased grass yield and the size of grass leaves and increased grass height more than clover height; it also increased the proportion of petiole relative to leaflet in the clover. Differences between varieties in response to interval between harvests were small but supported the view that medium large-leaved varieties can with advantage be defoliated rather less frequently than small-leaved ones. The adverse effect of applied N on clover appeared almost equally great with all four intervals between harvests and further research on this topic is suggested. Applied N increased grass height more than clover height and increased the number of grass tillers, the size of grass leaves and grass yield. The medium large-leaved varieties seemed more tolerant of applied N than the smaller varieties.

The rate of growth of Italian ryegrass in the spring was measured by recording dry-matter yield at weekly intervals over periods of 6 weeks. There were 3 levels of applied N: 25, 75 and 125 lb/ac. Yield increased fairly slowly in the first 2 weeks and level of applied N had little effect. In the final 4 weeks the grass grew at a faster and fairly constant rate, which was very much affected by the level of N. Grass receiving 25 lb N produced 2900 lb of dry matter/ac in 6 weeks, 75 lb N 4400 lb, and 125 lb N 4700 lb. By the end of the first week, % N and % nitrate-N were higher at the 75 lb than at the 25 lb N level and by the end of the second week they were higher at the 125 lb than at the 75 lb N level. The effect of weather factors and the question of predictability of yield are discussed.

Six varieties of white clover, each grown with perennial ryegrass, four intervals between cuts and two levels of applied nitrogen in all combinations, were compared in a field experiment during the first 27 months after sowing. Information about yields, crop fractions, heights and ryegrass tillers has been presented in an earlier paper (Wilman and Asiegbu, 1982). The present paper is concerned with the more detailed studies of white clover, which help to explain the yield results and contribute to the understanding of the response of this species to management when grown in competition with grass.Increasing the interval between harvests increased the length of clover stolon per unit area of ground and increased stolon diameter, petiole length, weight per leaf and number of leaves harvested as a proportion of the number present in the sward while only slightly reducing the rate of leaf emergence, helping to explain the positive effect of increasing the interval on clover yield noted in the earlier paper. During regrowth, successive leaves had longer petioles and the length of individual petioles increased beyond the stage at which the leaflets were fully opened. Weight per leaf in clover increased considerably from April to June and declined to below the April value by October. It was shown that weight per leaf can be greatly increased by increasing the interval between harvests without reducing the number of leaves harvested per unit area per year. The stolon length measurements provided some support for the view that medium large-leaved varieties of white clover can with advantage be defoliated rather less frequently than small-leaved varieties. Stolon length was less adversely affected by applied N in the medium large-than in the smallleaved varieties. The small-leaved varieties had thinner stolons than the medium large-leaved varieties but about twice the stolon length when no N was applied, and a relatively high proportion of leaves which escaped defoliation. The application of N reduced stolon diameter, increased petiole length and had little or no effect on weight per clover leaf.

The effects of 4 levels of applied nitrogen, ranging from nil to a maximum of 417 lb N/ acre/annum, in all combinations with 3 frequencies of defoliation, ranging from 2 to a maximum of 10 cuts per annum, on herbage production from a perennial ryegrass/ timothy/meadow fescue/white clover sward were measured. These treatments were operative for 2J years, and in a subsequent year the residual effect of cutting frequency was tested. Dry-matter yields of total herbage and of the clover fraction are quoted, together with N yields of total herbage. Yield response to N was higher than in some other experiments in the U.K. Cutting frequency had a very large effect and, in general, the longer the interval between cuts, the higher was the dry-matter (though not the N) yield. There was a marked interaction between cutting frequency and level of N: at the high cutting frequency, dry-matter yield increased linearly with increasing level of N; at the medium frequency, response tended to fall off at the highest level of N; at the low frequency, yield declined with increasing level of N beyond 139 lb N per acre per anum.

Leaf blades of perennial ryegrass {Lolium perenne) at five developmental stages, from unemerged to dead, were collected from cut swards grown under field conditions. Similarly, leaflets of white clover (Trifolium repens) at four developmental stages, from tightly folded to proximal to the youngest fully developed leaf, were collected. The leaves were analysed for N, P, K, Ca. Mg and Na.In perennial ryegrass, the concentration of N declined by about half and those of P and K declined by about two-thirds, as leaves aged. By contrast, the concentration of Ca increased more than fivefold and that of Mg about twofold as leaves aged. The concentration of Na initially increased more than twofold but then declined as leaves died. In white clover the concentration of N declined by about 20% as leaves aged, while the concentrations of P and K declined by about 50%; the concentration of Ca increased about fivefold, that of Mg declined by about 15% and that of Na increased by about 15%.The application of N increased the concentrations of N and K in ryegrass leaf blades at all stages of development. The application of P increased the concentrations of P and Na in the leaves of both species.