1. Physico‐chemical conditions and benthic macroinvertebrates were studied in two adjacent alpine streams in the Tyrolean Alps, Austria, for 2 years, and aquatic insect emergence was recorded for 1 year. 2. In the spring‐fed system, maximum discharge and increased concentrations of suspended solids, nitrate and particulate phosphorus occurred during snowmelt in June. In the glacier‐fed stream, high discharge and strong diel fluctuations in flow and concentrations of suspended solids created a harsh and unstable environment during summer. Glacial ablation, variation in groundwater inflow, and water inputs from tributaries draining calcareous rocks caused water chemistry to vary both seasonally and longitudinally in glacier‐fed Rotmoosache. 3. A total of 126 aquatic or semi‐aquatic invertebrate taxa were collected, 94 of which were found in the glacier‐fed stream and 120 in the spring‐fed stream. Chironomid abundance was 2–8 times and taxa richness 2–3 times lower in the glacier‐fed stream than in the spring‐fed stream, as was the number of chironomid taxa (72 versus 93 total). 4. These results broadly support the conceptual model by Milner & Petts (1994) concerning glacier‐fed stream systems. However, single samples and seasonal means showed relatively high invertebrate abundance and richness, especially during winter, indicating a considerable degree of spatial and temporal variability. 5. We suggest that the seasonal shifts from harsh environmental conditions in summer to less severe conditions in autumn and a rather constant environment in winter are an important factor affecting larval development, life‐history patterns and the maintenance of relatively high levels of diversity and productivity in glacier‐fed streams.
1. Alpine streams above the tree line are covered by snow for 6–9 months a year. However, winter dynamics in these streams are poorly known. The annual patterns of macroinvertebrate assemblages were studied in a glacial stream in the Austrian Alps, providing information on conditions under the snow. 2. Snow cover influenced water temperature, the content of benthic organic matter and insect development. Taxa richness and abundance of macroinvertebrates did not show a pronounced seasonal pattern. The duration of the autumn period with stable stream beds was important in determining the abundance and composition of the winter fauna. 3. There were significant differences in species composition between summer and winter. Two potential strategies in larval survival were evident: adaptation to the extreme abiotic conditions in summer (e.g. Diamesa spp.) or avoidance of these conditions and development during winter (e.g. Ephemeroptera and Plecoptera). 4. A comparison of a stream reach with continuous snow cover and a stream reach that remained open throughout winter showed that conditions under snow are suboptimal. At the open stream site, with higher water temperatures and greater food supply (benthic organic matter content), abundance and taxa richness was higher and larval growth was faster. Several taxa were found exclusively at this site. 5. Winter conditions did not provide an entirely homogeneous environment, abiotic conditions changed rapidly, especially at the onset of snowfall and at snowmelt. Continuous monitoring is necessary to recognize spatial and temporal heterogeneity in winter environments and the fauna of alpine streams.
High discharges at hydropower plants (HPP) may mask fishway attraction flows and, thereby, prevent fishes from locating and using fishways critical for their access to upstream spawning and rearing habitats. Existing methods for determining attraction flows are either based on simple guidelines (e.g., a proportion of HPP discharge) that cannot address the spatial and temporal complexity of tailrace flow patterns or complicated studies (e.g., combinations of detailed hydraulic and biological investigations) that are expensive and time-consuming. To bridge this gap, we present a new, intermediate approach to reliably determine attraction flows for technical fishways at small to medium-sized waterways (mean annual flow up to 400 m3/s). Fundamental to our approach is a design criterion that the attraction flow should maintain its integrity as it propagates downstream from the fishway entrance to beyond the highly turbulent zone characteristic of HPP tailraces to create a discernable migration corridor connecting the fishway entrance to the downstream river. To implement this criterion, we describe a set of equations to calculate the width of the entrance and the corresponding attraction discharge. Input data are usually easy to obtain and include geometrical and hydraulic parameters describing the target HPP and its tailrace. To confirm our approach, we compare model results to four sites at German waterways where the design of attraction flow was obtained by detailed experimental and numerical methods. The comparison shows good agreement supporting our approach as a useful, intermediate alternative for determining attraction flows that bridges the gap between simple guidelines and detailed hydraulic and biological investigations.
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