A large-scale hydrodynamical cosmological simulation, Horizon-AGN , is used to investigate the alignment between the spin of galaxies and the cosmic filaments above redshift 1.2. The analysis of more than 150 000 galaxies per time step in the redshift range 1.2 < z < 1.8 with morphological diversity shows that the spin of low-mass blue galaxies is preferentially aligned with their neighbouring filaments, while high-mass red galaxies tend to have a perpendicular spin. The reorientation of the spin of massive galaxies is provided by galaxy mergers, which are significant in their mass build-up. We find that the stellar mass transition from alignment to misalignment happens around 3 × 10 10 M ⊙ . Galaxies form in the vorticity-rich neighbourhood of filaments, and migrate towards the nodes of the cosmic web as they convert their orbital angular momentum into spin. The signature of this process can be traced to the properties of galaxies, as measured relative to the cosmic web. We argue that a strong source of feedback such as active galactic nuclei is mandatory to quench in situ star formation in massive galaxies and promote various morphologies. It allows mergers to play their key role by reducing post-merger gas inflows and, therefore, keeping spins misaligned with cosmic filaments.

The interplay between cosmic gas accretion on to galaxies and galaxy mergers drives the observed morphological diversity of galaxies. By comparing the state-of-the-art hydrodynamical cosmological simulations Horizon-AGN and Horizon-noAGN, we unambiguously identify the critical role of active galactic nuclei (AGN) in setting up the correct galaxy morphology for the massive end of the population. With AGN feedback, typical kinematic and morpho-metric properties of galaxy populations as well as the galaxy-halo mass relation are in much better agreement with observations. Only AGN feedback allows massive galaxies at the centre of groups and clusters to become ellipticals, while without AGN feedback those galaxies reform discs. It is the merger-enhanced AGN activity that is able to freeze the morphological type of the post-merger remnant by durably quenching its quiescent star formation. Hence morphology is shown to be driven not only by mass but also by the nature of cosmic accretion: at constant galaxy mass, ellipticals are galaxies that are mainly assembled through mergers, while discs are preferentially built from the in situ star formation fed by smooth cosmic gas infall.