Island evolution may be expected to involve fast initial morphological divergence followed by stasis. We tested this model using the dental phenotype of modern and ancient common voles (Microtus arvalis), introduced onto the Orkney archipelago (Scotland) from continental Europe some 5000 years ago. First, we investigated phenotypic divergence of Orkney and continental European populations and assessed climatic influences. Second, phenotypic differentiation among Orkney populations was tested against geography, time, and neutral genetic patterns. Finally, we examined evolutionary change along a time series for the Orkney Mainland. Molar gigantism and anterior‐lobe hypertrophy evolved rapidly in Orkney voles following introduction, without any transitional forms detected. Founder events and adaptation appear to explain this initial rapid evolution. Idiosyncrasy in dental features among different island populations of Orkney voles is also likely the result of local founder events following Neolithic translocation around the archipelago. However, against our initial expectations, a second marked phenotypic shift occurred between the 4th and 12th centuries AD, associated with increased pastoral farming and introduction of competitors (mice and rats) and terrestrial predators (foxes and cats). These results indicate that human agency can generate a more complex pattern of morphological evolution than might be expected in island rodents.

Scotland's First Settlers comprised a survey project to locate and examine sites relating to the earliest, Mesolithic, settlement of the Inner Sound, along the coastlands between Skye and the west coast of Scotland. Particular foci of interest included the existence and nature of midden sites, the use of rockshelters and caves, and the different types of lithic raw material in use. In addition, information relating to the human use of the area up to the present day was recorded. Fieldwork took place over five years between 1999 and 2004: the entire coastline of the Inner Sound together with its islands was walked; 129 new archaeological sites were recorded; 36 sites were shovel pitted; 44 test pitted; and one major excavation took place. Excavation at Sand has been particularly exciting as it has resulted in the analysis of a shell midden dating to the early-mid seventh millennium BC, the early Mesolithic of Scotland. This report comprises the results of survey and excavation work as well as detailed artefact reports, full information on ecofacts such as shell, and bone, and information on the development of the landscape and environment, including sea level change. Finally, the broad-scale coverage of the project has led to a number of discussion points that have much to offer further work, both within the area and further afield. Digital material associated with this project is available through Archaeology Data Service archive http://dx.doi.org/10.5284/1000285 Scotland's First Settlers

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BackgroundThe Mesolithic occupation of Scotland began soon after the end of the last glaciation, between 10,000 and 9000 years ago. Considerablc research has been undertaken in the past two decades (Mithen 2000; Pollard & Morrison 1996; Woodman 1989; Young 2000); much has been published, more is awaited, and work continues apace. Mesolithic sites occur throughout Scotland, though recent archaeological activity has been concentrated on the western seaboard.The coastal nature of much of the Scottish Mesolithic has long been recognized, although the contribution of inland sites is becoming more apparent. The relationship between shell middens and lithic scatters and the nature of the midden sites themselves are slowly becoming clearer (Bonsall 1996; Finlayson 1998), though the make-up of the material culture remains vague, as known early sites with preservation of organic materials are few and far between and specialists remain divided over their interpretation. More widely, it is generally recognized that the Mesolithic occurred during a time of dynamic enviromnental change although the impact on the human population remains to be documented.

This paper describes a small Mesolithic structure from the Cairngorm Mountains, Scotland. Excavations at Caochanan Ruadha identified a small oval structure (c. 3m x 2.2m) with a central fire setting, in an upland valley (c.540 m asl). The site was occupied at c. 8200 cal BP and demonstrates hunter-gatherer use of the uplands during a period of significant climatic deterioration. The interpretation of the structure is primarily based on the distribution of the lithic assemblage, as the heavily podsolised soils have left no trace of light structural features. The lithic assemblage is specialised, dominated by microlith fragments, and functional analysis has identified different uses of different areas inside the structure. The identification of small, specialised Mesolithic sites is unusual and this paper will discuss the evidence for the presence of the structure and its use, compare it to other Mesolithic structures in Britain and highlight some methodological implications.